Ancient European DNA of the Copper and Bronze Ages

Mitochondrial and Y-chromosome haplogroups extracted from Copper and Bronze Age human remains in Europe and related remains in Central Asia, arranged chronologically. Dates for bronze-making vary from region to region, so there is chronological overlap in the periods. Bell Beaker and Corded Ware / Single Grave / Battle Axe are classed here as Copper Age, though described as Neolithic in chronologies which exclude a Copper Age.For haplogroup nomenclature and aDNA issues see Introduction to ancient DNA

Culture

Country

Site and/or Individual

Sex

Date

Genome

Y-DNA

mtDNA

Source

pigmentation from genes
pigmentation surviving
usually approximate number of SNPs Haplogroup Additional information Haplogroup Additional information
Samara Eneolithic Russia Khvalynsk II, Volga River, Samara [I0122 / SVP 35] M 5200-4000 BCE 582950 R1b1 M415 H2a1 Mathieson 2015; Lazaridis 2016
Samara Eneolithic Russia Khvalynsk II, Volga River, Samara [I0433 / SVP 46] M 5200-4000 BCE 440667 R1a1 M459 U5a1i Mathieson 2015
Samara Eneolithic Russia Khvalynsk II, Volga River, Samara [I0434 / SVP 47] M 5200-4000 BC 62724 Q1a F2676 U4a2 or U4d Mathieson 2015
Copper Age tell Bulgaria Ivanovo [I2431 / 41] M 4725-4605 calBCE 738279 G2a2b2a1a1c1a

[CTS5990 / Z1903]

G2a2b2a1a1c1a:CTS4472:15663626A->G; G2a2b2a1a1c1a:CTS6763:17097462C->Ahet; G2a2b2a1a1c1a:CTS11388:23090368A->G; G2a2b2a1a1c1a:Z3423:19251438G->T; G2a2b2a1a:PF3346:24498136T->C; G2a2b2a1:PF3337:18023391G->T; G2a2b2a1:Y287:9381065G->T; G2a2b2a:CTS424:6744902C->T; G2a2b2a:CTS688:6953417T->C; G2a2b2a:CTS946:7100848A->G; G2a2b2a:CTS4454^:15655268C->T; G2a2b2a:CTS10366:19423576G->A; G2a2b2a:PF3329:8478026C->A; G2a2b2a:PF3342:21935606G->T; G2a2b2a:Z3243:21935550A->G; G2a2b:F2184:16512478G->A; G2a2b:F2535:17589788C->T; G2a2b:F3139:21151007T->C; G2a2b:Z3260:13671506G->T; G2a:CTS6026:16620480C->T; G2a:CTS6630:17022002C->T; G2a:CTS6753:17090976C->T; G2a:CTS9318:18819146T->A; G2a:F1975:15588776A->C; G2a:F3088:20813445G->A; G2a:F4086:7727677C->T; G2a:L31:14028148C->A; G2a:P15:23244026C->T; G2a:PF3141:23973594T->G; G2a:Z3240:10060449A->G; G2:CTS4264:15528792T->C; G2:CTS4413:15635425T->C; G2:CTS4703:15776024C->T; G2:CTS6316:16817402C->T; G2:CTS6742:17088129G->C; G2:CTS10089:19248446G->A; etc. N1b2 Mathieson 2017
Varna group Bulgaria Varna [ANI152 / VAR43] M 4683-4406 calBCE 26224 CT CT:M5578:6619341C->A; CT:M5738:17909505G->A; no data Mathieson 2017
Varna group Bulgaria Varna [ANI153 / VAR44] M 4551-4374 calBCE 23133 R1 R1:M306:22750583C->A; no data Mathieson 2017
Varna group Bulgaria Varna [ANI159-ANI181 / VAR117-I] M 4711-4530 calBCE 419597 G2a2b2b G2a2b2b:CTS5434:16271769A->G; G2a2b2b:F705:2659191T->C; G2a2b2b:F1175:8399197G->A; G2a2b2b:F1429:8841269G->T; G2a2b2b:F1581:9646643G->C; G2a2b2b:F1760:14323704T->C; G2a2b2b:F2419:17192221G->A; G2a2b2b:PF3359:7877472G->A; G2a2b2b:PF3404:17632283G->A; G2a2b2b:PF3413:19399866T->C; G2a2b2:CTS9957:19170699C->T; G2a2b:L30:15604899C->T; G2a:CTS9318:18819146T->A; G2a:CTS11463:23122426G->A; G2a:M3397:21605685G->C; G2a:M3408:22109159G->C; G2:CTS2406:14294068C->T; G2:CTS4264:15528792T->C; G2:CTS6742:17088129G->C; G2:F1294:8545324T->A; G2:F3220:21637589G->C; G2:F3344:22697266G->A; G2:M3579:21147058A->G; G:CTS282:2871867A->G; G:CTS827:7038432C->G; G:CTS995:7132348G->C; etc. no data Mathieson 2017
Varna group Bulgaria Varna [ANI160 / VAR127] M 4685-4499 calBCE 62189 G2 G2:CTS1900:14116322T->A; G2:CTS6742:17088129G->C; G:CTS34:2681740G->A; G:CTS1139:7231638A->G; G:F1131:8240725C->T; G:M3464:7537950C->T; G:S1435:13658486C->G; no data Mathieson 2017
Copper Age tell Bulgaria Smyadovo [I2430 / 40] M 4545-4450 calBCE 617793 R1b1a [L754] R1b1a:A702:10038192G->A; R1b1a:CTS8612:18109555C->A; R1b1a:FGC36:13822833G->T; R1b1a:FGC41:7900883C->A; R1b1a:L761:16773870A->G; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:L278:18914441C->T; R1:CTS2565:14366723C->T; R1:CTS5611:16394489T->G; R1:P236:17782178C->G; R:CTS6417:16882568T->C; etc. Mathieson 2017
Copper Age tell Bulgaria Smyadovo [I2181] M 4550-4455 calBCE 71542 R R:P280:21843090C->G; HV15 Mathieson 2017
Copper Age tell Bulgaria Sushina [I2426] M 4450-4264 calBCE 30449 CT CT:M5611:7778691C->T; K1 Mathieson 2017
Eneolithic (Stog/Post-Stog) Ukraine Revova, kurgan 3 [R3.19a] M 4274 ± 93 BC U4 7028T, 12308G, 16356C Nikitin 2017
Early Eneolithic Bulgaria Smyadovo [SMY3] 6500-5000 y. a. H rCRS in HV1 Wilde 2014
Early Eneolithic Bulgaria Smyadovo [SMY4] 6500-5000 y. a. H 456Y, 16304C Wilde 2014
Early Eneolithic Bulgaria Smyadovo [SMY9] 6500-5000 y. a. H 16261T Wilde 2014
Early Eneolithic Bulgaria Smyadovo [SMY11] 6500-5000 y. a. T2e 4216C, 4917G, 7028T, 11719A, 14766T, 16126C, 16153A, 16269G, 16294T, 16296T, reported as T Wilde 2014
Early Eneolithic Moldova Vinogradnoe [VIN1]
6500-5000 y. a. H rCRS in HV1 Wilde 2014
Ust-Tartas Russia Sopka [Ut 2] 4000-3000 BC U2e 16129C, 16189C Molodin 2012
Ust-Tartas Russia Sopka [Ut 12, Ut 16] 4000-3000 BC U2e 2 samples of 16129C, 16189C, 16246G Molodin 2012
Ust-Tartas Russia Sopka [Ut 3] 4000-3000 BC D 16223T, 16362C Molodin 2012
Ust-Tartas Russia Sopka [Ut 4, 7, 16 and 37] 4000-3000 BC C 4 samples of 16223T, 16298C, 16327T Molodin 2012
Ust-Tartas Russia Sopka [Ut 5 and 38] 4000-3500 BC A10* 2 samples of 663G, 16148T, 16223T, 16227C, 16290T, 16311C, 16319A Molodin 2012; Pilipenko 2015
Ust-Tartas Russia Sopka [Ut 9] 4000-3000 BC U5a1 16192T, 16256T, 16270T Molodin 2012
Ust-Tartas Russia Sopka [Ut 14] 4000-3000 BC U5a1 16192T, 16256T, 16270T, 16318 Molodin 2012
Ust-Tartas Russia Sopka [Ut 19] 4000-3000 BC U4 16356C Molodin 2012
Ust-Tartas Russia Sopka [Ut 31 and 32] 4000-3000 BC Z 2 samples of 16185T, 16223T, 16260T, 16298C, 16300G Molodin 2012
Ust-Tartas Russia Sopka [Ut 33 and 34] 4000-3000 BC Z 2 samples of 16185T, 16223T, 16260T, 16298C, 16298C Molodin 2012
Romania Urziceni [I4089 / URZI48] M 3761-3645 calBCE 784776 C1a2a C1a2a:S23981:22112268C->T; C1a2:V20:6845955G->A; C1a:CTS11043:22914979G->T; C1:CTS6773:17100606C->T; C1:F3393:23023974C->A; C:CTS182:2790145C->G; C:CTS2377:14281450G->A; etc. J1c Mathieson 2017
Maikop and Novosvobodnaya Russia Klady 3600-3000 BC U8, T2 and N1 Nedoluzhko 2015
Novosvobodnaya Russia Klady 3500 BC V7 T72C, A93G, C2515T, G9378A, C9541T, C11018T, A11720G, C11723T, G12851A, A14906G, A15302G, C15477T Nedoluzhko 2014
Late Eneolithic Bulgaria Durankulak [DUR1]
5500-4000 y. a. U52a2 7028T, 11719A, 12308G, 14766T, 14793G, 16114A 16192T 16256T 16270T 16294T, reported as U Wilde 2014
Late Eneolithic Ukraine Mayaki [MAJ8] 5500-4000 y. a. T2b 4216C, 4917G, 7028T, 11719A, 14766T, 16126C 16294T 16296T 16304C, reported as T Wilde 2014
Late Eneolithic Ukraine Mayaki [MAJ9] 5500-4000 y. a. W 7028T, 8251A, 11719A, 11947G, 12705Y, 14766T, 16223T 16292T Wilde 2014
Late Eneolithic Ukraine Molyukhov Bugor [MOB1] 5500-4,000 y. a. U5a1 7028T, 11719A, 12308G, 14766T, 14793G, 16192T 16256T 16270T 16399G, reported as U Wilde 2014
Late Eneolithic Ukraine Molyukhov Bugor [MOB 3] 5500-4000 y. a. U/K 7028T, 8251R, 11719A, 12308G, 14766T, 16192T 16256T 16270T 16399G Wilde 2014
Remedello Italy Remedello di Sotto [RISE487] M 3483-3107 BC 239530 I PF3661, Z16985, CTS4088, CTS6231, CTS8545, PF3780, L41;

Tagankin reads I2a1a1 (PF3983+ V2245/PF3976+ PF3972+ V1694/PF3969+ PF3865+ CTS10400/PF4045+ CTS3854/PF4007+ CTS2224/PF3998+ PF3863+ CTS7984/PF4029+, Y15473-, Y15462- , CTS11034/PF4099- PF4060- CTS6426/PF4094- L160/PF4013/S184- )

H2a 263G, 750G, 11203T, 15326G, 16519C Allentoft 2015; Mathieson 2015; additional info from Vladimir Tagankin
Globular Amphora

Poland Kierzkowo

[I2441 / 8.5] Son of I2440

M 3400-2800 BCE 510373 I2a2a1b I2a2a1b:CTS10057:19232160C->T; I2a2a1:CTS9183:18732197A->G; I2a2:L35:22725379C->A; I2a2:L181:19077754G->T; I2a2:P217:7628484C->T; I2:PF3664:8567995G->A; I:CTS1800:14073053G->A; I:CTS2536:14352669G->A; etc. K1b1a1 Mathieson 2017
Spain El Portalón, Sierra de Atapuerca [ATP 3] M 5466-5312 cal BP
K1a2b 12397G, 146T, 152T, 195T, 247G, 497T, 769G, 825T, 1018G, 1811G, 2758G, 2885T, 3480G, 3594C, 4104A, 4312C, 7146A, 7256C,
8005C, 8468C, 8655C, 8701A, 9055A, 9540T, 9698C, 10550G, 10664C, 10810T, 10873T, 10915T, 11025C, 11299C, 11467G, 11914G,  2308G, 12372A, 12705C, 13105A, 13276A, 13506C, 13650C, 14167T, 16093C, 16129G, 16187C, 16189T, 16223C, 16224C, 16230A, 16278C,
Günther 2015
Portugal Cova da Moura [CM 183*] 3400 BC H CRS, -7025 Afonso 2010
Portugal Cova da Moura [CM 187] 3400 BC X2b 16189C, 16223T, 16278T, +14465 Afonso 2010
Portugal Cova da Moura [CM 194 ] 3400 BC H 16111T, 16129A, +7025 Afonso 2010
Portugal Cova da Moura [CM 202] 3400 BC U5 or HV2 16217C, 16270A Afonso 2010
Bulgaria Beli Breyag [Bul6] M 3400-1600 BCE 370439 I2a2 I2a2:L37:17516123T->C; I2a2:L368:6931594C->Thet; I:CTS2193:14214481G->T; etc. - Mathieson 2017
Bulgaria Beli Breyag [Bul8] M 3400-1600 BCE 18337 I I:L758:8536868C->G; - Mathieson 2017
Italy Ötztal Alps [Otzi]
Brown eyes; brown hair; fair skin
M 3350-3100 BC 1,209,114 G2a1b2 L91 K1f 263G, 8137T, 1189C, 1811G, 9055A, 9698C, 14167T, 16224C, 16311C, 16362C, reported as K1ö.

Mitosearch: 4q6ja

Rollo 2006; Ermini 2008; Endicott 2009;
Keller 2012
Yamnaya Russia Lopatino I, Sok River, Samara [I0429 / SVP 38] M 3339-2917 BC 616,123 R1b1a2a2* (Z2103) Z2105+, L23+, L150+, M269+, L584- T2c1a2 Haak 2015; Mathieson 2015; Sergey Malyshev
Copper Age tell Bulgaria Smyadovo [I2176 / 12 Burial 20B] Silver hair ring M 3338-3025 calBCE I2a2a1b [CTS10057] I2a2a1b:CTS10100:19255890G->A; I2a2a:L34:7716262A->C; I2a2a:P223:16699334C->G; I2a2:L37:17516123T->C; I2a2:L368:6931594C->T; I2:L68:18700150C->T; I:CTS646:6926038T->A; I:CTS1800:14073053G->A; etc. U1a1 Mathieson 2017
Bulgaria Dzhulyunitsa [I2520] M 3336-3028 calBCE 795071 H2 H2:L279:6932824G->T; H2:L285:21869856C->T; H2:P96:14869743C->A; H:M2713:6855809G->A; H:M2896:16919982C->T; etc. H Mathieson 2017
Globular Amphora

Poland Kierzkowo

[I2405 / 8.2a]

M 3335-3020 calBCE

91505 I2a2 I2a2:P218:17493630T->G; I:CTS10941:22845794A->G; W5 Mathieson 2017
Yamnaya Russia Peshany V [RISE550] M 3334-2635 BC 476829 R1b1a2 [*R1b1a2a2] PF6399, M520, L773, PF6430, PF6434, PF6438, CTS8728, PF6497, PF6500;

*Malyshev reads Z8129/Y12537 (equivalent Z2103)

U5a1i 73G, 263G, 1438G, 2706G, 3197C, 9477A, 11467G, 11719A, 12308G, 12372A, 13617C, 14766T, 14793G, 14893G, 14971C, 15218G, 15326G, 16192T, 16256T, 16270T, 16399G Allentoft 2015; Mathieson 2015; *additional info from Sergey Malyshev
Afanasievo Russia River Kuyum [RISE508] F 3331-2935 calBCE (4442±29 BP) 322453 U5a1a1 73G, 263G, 750G, 1700C, 2706G, 3197C, 5495C, 9030T, 11467G, 12308G, 12372A, 13617C, 14793G, 15218G, 15326G, 15924G, 16256T, 16270T, 16399G Allentoft 2015
Copper Age tell Bulgaria Smyadovo [I2175 / 10 Burial 20A] Silver hair ring M 3328-3015 calBCE 423781 I2a2a1b1 [L701, L702] I2a2a1b1:L702:7629205C->T; I2a2a1b:CTS10057:19232160C->T; I2a2a1:CTS9183:18732197A->G; I2a2a:P220:24475669G->T; I2a2a:P221:8353707C->A; I2a2a:P222:18888200C->G; I2:M438:16638804A->G; I:CTS1800:14073053G->A; I:CTS4209:15479899T->A; etc. K1c1 Mathieson 2017
Afanasievo Russia River Kuyum [RISE507] F 3322-2923 BC 170,131 U5a1a1 73G, 263G, 750G, 1700C, 2706G, 3197C, 5495C, 9030T, 11467G, 12308G, 12372A, 13617C, 14793G, 15218G, 15326G, 15924G, 16256T, 16270T, 16399G Allentoft 2015
Yamnaya Russia Lopatino I, Sok River, Samara [I0439 / SVP 52] M 3305-2925 BC 273652 R1b1a2 PF6434, PF6475, L1353, M173+, M343 equivalent CTS910/ FGC66 /PF6244+; P297+, M269 equivalent CTS12478/ PF6529+, L51-, L584-, Y4370- M12147-, 14168106(A/G)+ U5a1a1 Haak 2015; Mathieson 2015; Sergey Malyshev
Yamnaya Russia Ishkinovka I, Eastern Orenburg, Pre-Ural steppe, Samara [I0370 / SVP 10] M 3300-2700 BC 712941 R1b1a2a2 [Z2103] CTS1078+, M269+, L150+, L320+ H13a1a1a Haak 2015; Mathieson 2015; Lazaridis 2016; Sergey Malychev
Yamnaya Russia Lopatino II, Sok River, Samara [I0443 / SVP 57] M 3300-2700 BC 1025251 R1b1a2a* [L23] L49+, L23+, PF6399+, L150+, L1353+, PF6509+, M269+, CTS12478+, L51-, Z2105- W3a1a Haak 2015; Mathieson 2015; Sergey Malychev
Yamnaya Ukraine Shevchenko [I3141] F 3300-2700 BC 45365 H15b1 Mathieson 2017
Yamnaya Ukraine Shevchenko [I2105] F 3300-2700 BC 802473 T1a1 Mathieson 2017
Yamnaya Russia Kutuluk I, Kutuluk River, Samara [I0444 / SVP 58] M 3300-2700 BC 551461 R1b1a2a2 [Z2103] CTS1078/Z2103+, L150+, M415+ H6a1b Haak 2015; Mathieson 2015; Sergey Malychev
Portugal Dólmen do Ansiăo [DEA 50*] 3300 BC H or U4 16356C Afonso 2010
Spain El Portalón, Sierra de Atapuerca [ATP7] F 5295-4894 cal BP J1c1b1 146T, 152T, 185A, 195T, 228A, 247G, 295T, 482C, 769G, 825T, 1018G, 2758G, 2885T, 3010A, 3394C, 3594C, 4104A, 4312C, 5773A, 7146A, 7184G, 7256C, 7521G, 8468C, 8655C, 8701A, 9540T, 10688G, 10810T, 10873T, 10915T, 11914G, 12705C, 13105A, 13276A, 13506C, 13650C, 14798C, 16069T, 16126C, 16129G, 16187C, 16189T, 16223C, 16230A, 16278C, 16311T, 4769A, 6156T, 6169T, 6172T, 6173T, 16519T Günther 2015
Spain El Portalón, Sierra de Atapuerca [ATP16] F 3261-2916 calBC (4400±30, Beta-368289) 839051 X2c 146T, 152T, 153G, 225A, 227G, 247G, 769G, 825T, 1018G, 1719A, 2758G, 2885T, 3594C, 4104A, 4312C, 6371T, 7146A, 7256C, 7521G, 8468C, 8655C, 8701A, 9540T, 10398A, 10664C, 10688G, 10810T, 10873T, 10915T, 11914G, 13105A, 13276A, 13506C, 13650C, 13966G, 14470C, 16129G, 16187C, 16230A, 16255A, 16311T, 15314A, 16207G Günther 2015
Globular Amphora

Poland Kierzkowo

[I2440 / 7.6]

Father to I2441

M 3100-2900 BCE

189493 I2 I2:L68:18700150C->T; I:CTS7469:17497181C->A; I:FGC2412:21689728A->G; I:FGC2413:8262092C->T; I:FI2:8382265C->G; I:FI4:8873160G->T; I:L578:8267857G->A; I:PF3640:7681156T->A; I:PF3641:7688470T->C; H1b Mathieson 2017
Yamnaya Ukraine Ozera, Kurgan 18, grave 14 [I1917 / OAE-99 / Yamna1]

F 3095-2915 calBCE 602225

[Genetically mixture EEF and Samara Yamnaya]

R0a1 Mathieson 2017
Ezero Bulgaria Sabrano [Bul10 / Sabrano, Grave 1, Individual 1D] F 3090-2924 calBCE 66247 - Mathieson 2017
Yamnaya Russia Lopatino I, Sok River, Samara [I0357 / SVP 5] F 3090-2910 BC 544,122 W6c Haak 2015; Mathieson 2015
Corded Ware Latvia Zvejnieki [I4629 / ZVEJ28/ LN1] F 3089-2676 calBCE 251797 U5a1b Jones 2017; Mathieson 2017
Yamnaya Russia Luzhki I, Samara River, Samara [I0438 / SVP 50] M 3021-2635 BC 579,356 R1b1a2a2* [Z2103] Z2105+, L23+, L320+, L584-, CTS7822- U5a1a1 Haak 2015; ; Mathieson 2015; Sergey Malychev
Yamnaya Bulgaria Mednikarovo [Bul4 / Barrow 2, grave 1] M 3012-2900 calBCE

372047 [Genetically mixture EEF, EHG, and Samara Yamnaya] I2a2a1b1b [L699] I2a2a1b1b:L699:2663920A->G; I2a2a1:CTS9183:18732197A->G; I2a2a:P221:8353707C->A; I2a2:L37:17516123T->C; I2a2:L181:19077754G->T; I2a2:P218:17493630T->G; I:CTS2387:14286853T->C; etc. - Mathieson 2017
Yamnaya Russia Kurmanaevskii III, Buzuluk, Samara [I0441 / SVP 54] F 3010-2622 BC 104,430 H2b Haak 2015; Mathieson 2015; Lazaridis 2016
Portugal Paimogo [PM 13297] 3000 BC H or HV* CRS, -12308, -11719 Afonso 2010
Portugal Paimogo [PM 13514] 3000 BC HV* or L3*/N* 16223T, 16278T, 16311C Afonso 2010
Portugal Paimogo [PM 13622] 3000 BC U CRS, +7025 +12308 Afonso 2010
Portugal Paimogo [PM 14638] 3000 BC U5b2* 16192T, 16217C, 16270A, +12308 Afonso 2010
Portugal Paimogo [PM 14714] 3000 BC H CRS, -7025, -12308, -11719 Afonso 2010
Portugal Paimogo [PM 15079] 3000 BC U4 16356C, +7025, +12308 Afonso 2010
Portugal Paimogo [PM 15986] 3000 BC H 16270T, -7025, -12308, -11719 Afonso 2010
Portugal Paimogo [PM 17334] 3000 BC H 16147T, -12308, -11719 Afonso 2010
Portugal Paimogo [PM 17480] 3000 BC H, HV* or R0 CRS, -12308 Afonso 2010
Portugal Paimogo [PMII 527, 542] 3000 BC H 2 samples 16183C, 16189C, -7025 Afonso 2010
Portugal Paimogo [PMII 556] 3000 BC H CRS, -7025 Afonso 2010
Portugal Paimogo [PMII 591] 3000 BC H 16292G, -7025, -11719 Afonso 2010
Portugal Monte Canelas [MCI 169.23*] 3000 BC H, HV* or R0 CRS, -12308 Afonso 2010
Portugal Monte Canelas [MCI 176.1] 3000 BC H 16294T, 16304C, -7025, -12308, -11719 Afonso 2010
Portugal Monte Canelas [MCI 206.4*] 3000 BC H CRS, -7025. Afonso 2010
Portugal Monte Canelas [MCI282.3.1] 3000 BC H or V 16256T, -12308 Afonso 2010
Portugal Monte Canelas [MCI 288.5] 3000 BC H 16256T, -7025 Afonso 2010
Portugal Monte Canelas [MCI 337*] 3000 BC H 16356C, -7025 Afonso 2010
Portugal Monte Canelas [MCI 384.11*] 3000 BC H 16311C, 16354T, -12308 Afonso 2010
Portugal Monte Canelas [MCI 386.37 ] 3000 BC U5a1 16256T, 16270T, 16274A Afonso 2010
Portugal Monte Canelas [MCI 435.3] 3000 BC H 16093C, -7025 Afonso 2010
Yamnaya Bulgaria Benkovski [BEN3] 5000-4500 y.a. H 4937C, 8935T, 13269G, 16188T Wilde 2014
Yamnaya Russia Kalinovka I [KAL1] 5000-4500 y.a. N1a 669C, 7028T, 10398G, 11719A, 12705T, 14766T, 16147G 16172C 16223T 16248T 16344T 16355T Wilde 2014
Yamnaya Russia Kalinovka I [KAL2] 5000-4500 y.a. H Wilde 2014
Yamnaya Ukraine Mayaki [MAJ3] 5000-4500 y.a. ? 7028T, 11719A, 14766T, 16256T 16270T 16399G Wilde 2014
Yamnaya Ukraine Mayaki [MAJ4] 5000-4500 y.a. U5 4194Y, 7028T, 11719A, 12308G, 13637G, 14766T, 16192T 16311C, reported as U Wilde 2014
Yamnaya Ukraine Mayaki [MAJ5] 5000-4500 y.a. X 6371T, 7028T, 11719A, 12705T, 14766T, 16136C 16189C 16223T 16278T Wilde 2014
Yamnaya Russia Nikolaevka III [NIK1] 5000-4500 y.a. T1a 4216C, 4917G, 7028T, 11719A, 12346T, 14766T, 16163G 16186T 16189C 16294T, reported as T Wilde 2014
Yamnaya Russia Nikolaevka III [NIK7] 5000-4500 y.a. H 13680T, rCRS in HV1 Wilde 2014
Yamnaya Russia Olennii [OLE1] 5000-4500 y.a. T 4126C, 4917G, 7028T, 11719A, 14766T, 16126C 16292T 16294T Wilde 2014
Yamnaya Russia Olennii [OLE7]
5000-4500 y.a. J 4126C, 7028T, 10398G, 11719A, 13708A, 14766T, 14769G, 16048A 16069T 16126C 16193T Wilde 2014
Yamnaya Bulgaria Ovchartsi [OVI2]
5000-4500 y.a.

K 7028T, 10398G, 11719A, 12308G, 14766T, 14798C, 16224C 16290T 16311C Wilde 2014
Yamnaya Bulgaria Ovchartsi [OVI3]
5000-4500 y.a.

U/K 11719A, 12308G, 14766T Wilde 2014
Yamnaya Russia Peschanyi [PEJ1]
5000-4500 y.a.

U5a1 7028T, 11719A, 12308G, 14766T, 14793G, 16192T 16256T 16270T 16399G Wilde 2014
Yamnaya Ukraine Pestchanka [PES7]
5000-4500 y.a.

H 481T, 16042A 16209C Wilde 2014
Yamnaya Russia Podlesnyj [POD1]
5000-4500 y.a.

W 7028T, 8251A, 8994A, 11719A, 11947G, 12705T, 14766T, 16192T 16223T 16292T 16325C Wilde 2014
Yamnaya Russia Podlesnyj [POD2]
5000-4500 y.a.

T 4216C, 4917G, 7028T, 11719A, 14766T, 16126C 16292T 16294T Wilde 2014
Yamnaya Bulgaria Golyamata Mogila, Popovo [POP1]
5000-4500 y.a.

T2a1b1a 4216C, 4917G, 7028T, 9117C, 11719A, 14766T, 16126C 16294T 16296T 16324C, reported as T Wilde 2014
Yamnaya Bulgaria Golyamata Mogila, Popovo [POP3]
5000-4500 y.a.

U2e1a 7028T, 10876G, 11719A, 11732C, 12308G, 13734C, 14766T, 16051G 16129C 16362C, reported as U Wilde 2014
Yamnaya Bulgaria Golyamata Mogila, Popovo [POP4]
5000-4500 y.a.

U5a1 7028T, 11719A, 12308G, 14766T, 14793G, 16256T 16270T 16399G, reported as U Wilde 2014
Yamnaya Bulgaria Riltsi [RIL3]
5000-4500 y.a.

K 7028T, 10398G, 11719A, 12308G, 14766T, 14798C, 16192T 16224C 16311C Wilde 2014
Yamnaya Ukraine Kirovograd Sugokleya [SUG2]
5000-4500 y.a.

I 4529T, 7028T, 8251A, 8260C, 10398G, 11719A, 12705T, 14766T, 16129A 16172C 16223T 16311C 16391A Wilde 2014
Yamnaya Ukraine Kirovograd Sugokleya [SUG6]
5000-4500 y.a.

H 7028Y, 16362C Wilde 2014
Yamnaya Ukraine Kirovograd Sugokleya [SUG7]
5000-4500 y.a.

H rCRS in HV1 Wilde 2014
Yamnaya Ukraine Kirovograd Sugokleya [SUG8]
5000-4500 y.a.

H 3010R, 14766T, rCRS in HV1 Wilde 2014
Yamnaya Moldova Tetcani [TET2]
5000-4500 y.a.

U 7028T, 11719A, 12308G, 14766T, 16134T 16356C Wilde 2014
Yamnaya Ukraine Vinogradnoe [VIN2, VIN5]
5000-4500 y.a.

T1a 2 samples of 4216C, 4917G, 7028T, 11719A, 14766T, 16126C 16163G 16186T 16189C 16294T, reported as T Wilde 2014
Yamnaya Ukraine Vinogradnoe [VIN12]
5000-4500 y.a.

T1 4216C, 4917G, 7028T, 11719A, 14766T, 16126C 16163G 16294T, reported as T Wilde 2014

Spain El Portalón, Sierra de Atapuerca [ATP17] M 4957-4821 cal BP

H3 73A, 146T, 152T, 195T, 247G, 769G, 825T, 1018G, 2706A, 2758G, 2885T, 3594C, 4104A, 4312C, 6776C, 7256C, 8468C, 8655C, 8701A, 9540T, 10398A, 10664C, 10688G, 10810T,
10873T, 10915T, 11719G, 11914G, 12705C, 13105A, 13276A,
13506C, 13650C, 14766C, 16129G, 16187C, 16189T, 16223C, 16230A, 16278C, 16311T, 4769A, 7702A, 7729G,
8573A, 9182A, 16093C
Günther 2015
Baltic LN Lithuania Plinkaigalis [242] F 3260-2630 cal BCE 861862 W6a Mittnik 2017
Poltavka outlier Russia Potapovka I, Sok River, Samara [I0432 / SVP 42] M 2925-2536 BC 648,053 R1a1a1b2a Z94 U5a1c Mathieson 2015
Spain Camino del Molino, Caravaca de la Cruz, Murcia [I0456 / Cmol140] F 2920–2340 BCE 28463 U5b2a1a Olalde 2017
Spain Camino del Molino, Caravaca de la Cruz, Murcia [I0457 / Cmol165b] M 2920–2340 BCE 19814 CF CF:P143:14197867G->A J1c1 Olalde 2017

Spain El Portalón, Sierra de Atapuerca [ATP12-1420; Matojo] M 2919 ± 18 calBC (4300±30, Beta-368295) 102,497 I2a2a
H3c 73A, 146T, 152T, 195T, 247G, 769G, 825T, 1018G, 2706A,
2758G, 2885T, 3594C, 4104A, 4312C, 6776C, 7028C, 7146A,
7256C, 7521G, 8468C, 8655C, 8701A, 9540T, 10398A, 10664C, 10688G, 10810T, 10873T, 10915T, 11719G, 11914G, 12705C, 12957C, 13105A, 13276A, 13506C, 13650C, 14766C,
16129G, 16187C, 16189T, 16223C, 16230A, 16278C, 16311T, 7493T, 8860A, 16399G
Günther 2015
Yamnaya Russia Ekaterinovka, Southern Steppe, Samara [I0231/SVP 3] M 2910-2875 BC 1,099,877 R1b1a2a2 + KMS75 CTS1078+, Z2105+, S20902+, M343+, L754/ PF6269/ YSC22+, P297 equivalent FGC57+, M269+ (+ same level PF6399+, PF6434+, PF6482+, , L150.1+, L265+), L23+, CTS1078/ Z2103+ (+ same level Z2105+), L584-, KMS75+ U4a1a or U4a1d
Haak 2015; Mathieson 2015; Sergey Malyshev
Afanasievo Russia Bateni [RISE511] F 2909-2679 BC 1,102,979



J2a2a 73G, 150T, 195C, 235G, 263G, 295T, 489C, 709A, 1438G, 2706G, 3107T, 6671C, 8386T, 10398G, 10499G, 11002G, 11251G, 11377A, 11719A, 12570G, 12612G, 13708A, 14766T, 15257A, 15326G, 15452A, 15679G, 16069T, 16126C Allentoft 2015
Remedello Italy Remedello di Sotto [RISE489] M 2908-2578 BC 510,496 I [*I2a1a1] CTS1301, FGC2413, PF3665, Z16985, CTS4848, CTS5946, CTS6231, CTS6343, CTS9264, PF3811, PF3815;

[*Tagankin reads I2a1a1 (Y3995+ PF4069+ PF4065+ PF4054+ PF4048+ PF4046+ PF3983+ V2245/PF3976+ PF3875+ PF3848+ PF3832/S6104+ CTS8644/PF4033+ CTS575/PF3952+ PF4074+ PF4055+ PF4049+ PF4014+ PF3977/S6233+ PF3874+ PF3863+ CTS11548/PF3877+ V3296/CTS8968/PF4036+ CTS7964/PF3767+ CTS6027+ CTS3353/PF3866+ CTS3276/PF3713+ CTS3128/PF4005+ CTS1581/PF3993+ CTS799/PF3954+, PF4096- PF4066- CTS4506/PF4010- I-PF4188: PF4195- PF4189- PF4188-, Y15654- Z27397+ Y15580-) ]

X2c1 73G, 153G, 195C, 225A, 227G, 263G, 750G, 1438G, 1719A, 2706G, 6371T, 8705C, 11719A, 12705T, 13966G, 14470C, 14766T, 15326G, 16223T, 16255A, 16278T, 16519C Allentoft 2015; Mathieson 2015; *additional info from Vladimir Tagankin
Bulgaria Dzhulyunitsa [I2510]

M 2906-2710 calBCE

821681 G2a2a1a2 G2a2a1a2:L91:21645555G->C; G2a2a1a2:PF3233:8895958T->C; G2a2a1:PF3155:14006343T->C; G2a2a1:PF3170:18090604G->A; G2a2a:PF3147:7738069G->A; G2a2a:PF3150:8476569T->C; G2a2a:PF3151:9785736A->G; G2a2a:PF3159:14815695C->G; G2a2a:PF3175:18962113C->T; G2a2a:PF3181:21808944C->A; G2a2a:PF3184:22576860C->T; G2a2a:PF3185:22894488C->T; G2a2a:Z6178:7245721G->A; G2a2:CTS4367:15615340C->G; G2a:CTS6753:17090976C->T; G2a:CTS11463:23122426G->A; G2a:F2529:17571517A->G; G2a:F3088:20813445G->A; G2a:F4086:7727677C->T; G2a:L31:14028148C->A; G2a:M3397:21605685G->C; G2a:P15:23244026C->T; G2a:PF3141:23973594T->G; G2a:Z3240:10060449A->G; G2:CTS2406:14294068C->T; G2:CTS4136:15421357G->A; G2:CTS4413:15635425T->C; G2:CTS4703:15776024C->T; G2:CTS5666:16424034C->T; G2:CTS6316:16817402C->T; G2:CTS7662:17597715C->T; G2:CTS10089:19248446G->A; G2:F1239:8482393C->T; G2:F1294:8545324T->A; G2:F1393:8719593G->A; G2:F1647:9907842G->T; G2:F2319:16903051A->T; G2:F3198:21401188G->T; G2:F3220:21637589G->C; G2:F3344:22697266G->A; G2:F3536:23768744C->T; G2:L89:7978725C->T; G2:M3465:7571775G->T; G2:M3579:21147058A->G; G2:P287:22072097G->T; G2:PF2909:13205148A->G; G2:PF3119:23739606G->T; G:CTS34:2681740G->A; G:CTS175:2785630A->T; etc. H4a1 Mathieson 2017
Baden Hungary Apc-Berekalja I [I1497 / CO1] F 2900-2700 BC 843,595

H 263G, 750G, 1438G, 4769G, 8860G, 15326G, 16519C*
Gamba 2014; Mathieson 2015; Lazaridis 2016
Spain Camino del Molino, Caravaca de la Cruz, Murcia [I0455 / Cmol123] M 2900–2670 calBCE 46870 I I:CTS7593:17548890G->A; I:L772:15615533C->A H1 Olalde 2017

Spain El Mirador, Burgos [I1282 / MIR 19] M
2900-2346 BCE

69,288 CTS4209, CTS4273
H3 Gómez-Sánchez 2014 ; Mathieson 2015; Lazaridis 2016
Globular Amphora

Ukraine Ilyatka [ILK001] M 2899-2706 calBCE 926232 I2a2a1b I2a2a1b:CTS10057:19232160C->T; I2a2a1b:CTS10100:19255890G->A; I2a2a1:CTS9183:18732197A->Ghet; I2a2a:L36:17570599C->T; I2a2a:L59:7113556C->T; I2a2a:P221:8353707C->A; I2a2a:P222:18888200C->G; I2a2a:P223:16699334C->G; I2a2:L35:22725379C->A; I2a2:L37:17516123T->C; I2a2:L181:19077754G->T; I2a2:L368:6931594C->T; I2a2:P216:13992338C->G; I2a2:P217:7628484C->T; I2a2:P218:17493630T->G; I2a:L460:7879415A->C; I2:L68:18700150C->T; I2:PF3664:8567995G->A; I:CTS674:6943522C->T; I:CTS1301:7321418C->T; I:CTS1800:14073053G->A; etc. J1c3 Mathieson 2017
Globular Amphora

Ukraine Ilyatka [ILK002] M 2890-2694 calBCE I2a2a1b2 [Z161] I2a2a1b2:Z161:2696497C->G; I2a2a1b:CTS10057:19232160C->T; I2a2a1:CTS9183:18732197A->G; I2a2a:L34:7716262A->C; I2a2a:P220:24475669G->T; I2a2a:P221:8353707C->A; I2a2a:P222:18888200C->G; I2a2:L35:22725379C->A; I2a2:L37:17516123T->C; I2a2:L181:19077754G->T; I2a2:L368:6931594C->T; I2a2:P216:13992338C->G; I2a2:P217:7628484C->T; I2a2:P218:17493630T->G; I2a:L460:7879415A->C; I2:L68:18700150C->T; I2:PF3664:8567995G->A; I:CTS88:2723755G->A; I:CTS646:6926038T->A; I:CTS674:6943522C->T; I:CTS1800:14073053G->A; etc. T2b Mathieson 2017

Spain El Portalón, Sierra de Atapuerca [ATP2] M 2899-2678 calBCE (4210±30, Beta-386394) 1205481

H2

U5b3 146T, 150T, 152T, 228A, 247G, 769G, 825T, 1018G, 2758G,
2885T, 3197C, 3594C, 4104A, 4312C, 7146A, 7226A, 7256C,
7521G, 8468C, 8655C, 8701A, 9477A, 9540T, 10398A,
10664C, 10688G, 10810T, 10873T, 10915T, 11467G, 11914G,
12308G, 12372A, 12705C, 13105A, 13276A, 13506C, 13617C,
13650C, 14182C, 16129G, 16187C, 16189T, 16192T, 16223C,
16230A, 16270T, 16278C, 16304C, 16311T, 6079T, 8860A, 16519T
Günther 2015; Lazaridis 2016
Afanasievo Russia Bateni [RISE509] F 2887-2677 BC 703,520

T2c1a2 73G, 263G, 709A, 1438G, 2706G, 3107T, 4917G, 5262A, 5817T, 6261A, 8697A, 10463C, 10822T, 11251G, 11719A, 11812G, 13973T, 14233G, 14766T, 14905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16292T, 16294T, 16519C Allentoft 2015
Yamnaya Russia Temrta IV [RISE546] M
167261

R1b1a2

PF6482 U5a1d2b 73G, 263G, 750G, 1438G, 3107T, 3197C, 4924C, 9477A, 10858C, 11467G, 11719A, 12308G, 12372A, 13617C, 14110C, 14766T, 14793G, 15217A, 15218G, 15326G, 16192T, 16256T, 16270T, 16304C, 16399G Allentoft 2015; Mathieson 2015
Yamnaya Russia Temrta IV [RISE547] M 2887-2634 BC 628581 R1b1a2 L482, L483, PF6438, L150.1, F1794, CTS8728, L1353, PF6495, PF6497;

Malyshev reads R1b1a2a2 (CTS9416) + Z2106 downstream on Y-Full tree

T2a1a 73G, 263G, 709A, 750G, 1438G, 2706G, 2850C, 3107T, 4917G, 7022C, 7028T, 8697A, 10463C, 11251G, 11719A, 11812G, 13368A, 13907G, 13965C, 14233G, 14687G, 14766T, 14905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16294T, 16296T, 16519C Allentoft 2015; Mathieson 2015; additional info from Sergey Malyshev
Yamnaya Russia Temrta IV [RISE548] M
733271 R1b1a2a2 Z2105 U4 73G, 195C, 263G, 499A, 750G, 1438G, 1811G, 3107T, 11332T, 11467G, 11719A, 12308G, 12372A, 14620T, 14766T, 15326G, 15693C, 16356C, 16519C Allentoft 2015; Mathieson 2015
Yamnaya Russia Sukhaya Termista I [RISE240] F 2880-2632 BC 212900

U5a1d1 73G, 263G, 750G, 1438G, 2706G, 3027C, 3107T, 3197C, 5263T, 9477A, 11467G, 11719A, 12308G, 12372A, 13002A, 13617C, 14766T, 14793G, 15218G, 15326G, 16192T, 16256T, 16270T, 16399G Allentoft 2015

Spain El Mirador, Burgos [I1271/MIR 1] F
2880-2630 BC
K1a 16294T 16296T 16324C Gómez-Sánchez 2014; Mathieson 2015
Spain El Mirador, Burgos [I1272/MIR 2] F 2880-2630 BC K1b1a1 16189C 16223T 16278T 16352C Gómez-Sánchez 2014 ; Mathieson 2015
Spain El Mirador, Burgos [MIR 3]
2880-2630 BC
H CRS Gómez-Sánchez 2014 ; Mathieson 2015

Spain El Mirador, Burgos [MIR 4]
2880-2630 BC

T2 16126C 16294T 16296T 16324C Gómez-Sánchez 2014 ; Mathieson 2015

Spain El Mirador, Burgos [I0581/MIR 5 and 6] M
2880-2630 BC I2a2a1 CTS616
X2b 189C 223T 278T (358T) Gómez-Sánchez 2014 ; Mathieson 2015

Spain El Mirador, Burgos [MIR 9]
2880-2630 BC

X2 16189C 16223T 16278T Gómez-Sánchez 2014

Spain El Mirador, Burgos [MIR 8, 15]
2880-2630 BC

T2b 2 samples 16126C 16294T 16296T 16304C Gómez-Sánchez 2014

Spain El Mirador, Burgos [MIR 10]
2880-2630 BC

K 16224C 16256T 16311C Gómez-Sánchez 2014

Spain El Mirador, Burgos [I1274 / MIR 11] M 2880-2630 BC 156079 I2a2
L181
H3 Gómez-Sánchez 2014 ; Mathieson 2015

Spain El Mirador, Burgos [MIR 12]
2880-2630 BC

K 16224C 16311C Gómez-Sánchez 2014

Spain El Mirador, Burgos [I1276 / MIR 13] F
2880-2630 BC

H3c3 16278T Gómez-Sánchez 2014 ; Mathieson 2015

Spain El Mirador, Burgos [MIR 16]
2880-2630 BC

K 16093C 16224T 16311C Gómez-Sánchez 2014 ;

Spain El Mirador, Burgos [I1280 / MIR 17] F 2880-2630 BC 208,463
J1c1 Mathieson 2015

Spain El Mirador, Burgos [MIR 20]
2880-2630 BC

J1+16193 / J2b 16069T 16126C 16193T 16274A 16278T Gómez-Sánchez 2014

Spain El Mirador, Burgos [I1284 / MIR 21] M 2880-2630 BC 125,450 I
L758, FGC2417, CTS5650 H3 Mathieson 2015

Spain El Mirador, Burgos [I1300 / MIR 22] F 2880-2630 BC 807,280

K1a2a 16224C 16311C Gómez-Sánchez 2014 ; Mathieson 2015

Spain El Mirador, Burgos [I1302 / MIR 24] M
2880-2630 BC 71,920 G2a2b2b F1760 J2b1a3 Mathieson 2015

Spain El Mirador, Burgos [I1303 / MIR 25] M 2880-2630 BC 468,021 I2a1a1
L158
U3a1 16343G Gómez-Sánchez 2014 ; Mathieson 2015

Spain El Mirador, Burgos [I1314/ MIR 26] M
2880-2630 BC 224117 G2a
PF3141
J2a1a1 16069T 16126C 16145A 16231C 16261T Gómez-Sánchez 2014
Corded Ware Germany Tiefbrunn [RISE434] M 2880-2630 BC 111108

P1 P239 U4 73G, 263G, 499A, 750G, 1438G, 1811G, 2706G, 3107T, 11332T, 11467G, 11719A, 12308G, 12372A, 14620T, 14766T, 15326G, 15693C, 16356C, 16519C Allentoft 2015; Mathieson 2015
Globular Amphora

Poland Kierzkowo

[I2403 / 3.4]

M 2870-2575 calBCE 290049 I2a2 I2a2:L181:19077754G->T; I2a2:P218:17493630T->G; I:CTS6265:16780748C->G; I:FI3:8485677C->A; I:L503:21359407C->G; I:L578:8267857G->A; U5b2b1 Mathieson 2017
Corded Ware Germany Tiefbrunn [RISE436] M 2868-2580 BC 63030 CT PF228, CTS3662, Y1528, CTS9948, Y1580 U5b1c2 73G, 150T, 516T, 2706G, 11719A, 12372A, 12705T, 13954T, 14766T, 15301A, 15326G, 16270T, 16311C Allentoft 2015; Mathieson 2015
Ukraine Katarzhyno, kurgan 2 main burial [K2.1] F 2866 ± 145 BC U5 12308G, 16241G, 16270T Nikitin 2017
Corded Ware Poland Oblaczkowo [RISE1] M 2865-2578 BC R1b R1b1 (L1345+) K1b1a1 73G, 152C, 263G, 709A, 1438G, 1811G, 2706G, 3480G, 3967A, 9962A, 10289G, 10550G, 11299C, 11467G, 11719A, 11923G, 12372A, 13967T, 14766T, 14798C, 15257A, 15326G, 16093C, 16224C, 16463G, 16519C Allentoft 2015; Y-DNA personal communication from author + additional info from Richard Rocca and Sergey Malyshev, citing Karmin 2015 for status of the SNP.

Spain El Mirador, Burgos [I1281 / MIR 18] F 2865-2575 calBCE (4110±30 BP) 646,025

H1t Gómez-Sánchez 2014 ; Mathieson 2015; Lazaridis 2016
Corded Ware Germany Tiefbrunn [RISE435] F 2863-2498 BC 50,680



J1b1a1 73G,242T, 263G, 295T, 462T, 489C, 750G, 1438G, 2158C, 2706G, 3010A, 3107T, 5460A, 7028T, 8269A, 8557A, 10398G, 11251G, 11719A, 12007A, 12612G, 13708A, 13879C, 14766T, 15326G, 15452A, 16069T, 16126C, 16145A, 16172C, 16222T, 16261T Allentoft 2015
Baltic LN Lithuania Plinkaigalis [241] F 2860-2410 cal BCE 190225 I2 Mittnik 2017

Hungary Lánycsók, Csata-alja [M6-116.8]
2860-2620 BC R1b M343+ T2b23 16126c 16147t 16294t 16296t 16297c 16304c 16362c Szécsényi-Nagy 2015 thesis

Hungary Lánycsók, Csata-alja [M6-116.10]
2860-2620 BC I2a2a M223+ H5 16304c Szécsényi-Nagy 2015 thesis
Early Bronze Age Russia Stalingrad Quarry [RISE555] M 2857-2497 BC 232,118

R1b1a2

[* R1b1a2a2]

PF6399, PF6409, PF6434, CTS623; [*Malyshev reads R1b1a2a2 (CTS7340/Z2107) + downstream Z2106 on YFull tree] N1a1a+152 73G, 152C, 199C, 204C, 263G, 669C, 1438G, 1719A, 2702A, 2706G, 3107T, 5315G, 8452G, 10238C, 10398G, 11476T, 11719A, 12501A, 12705T, 13780G, 14766T, 15043A, 15326G, 16147G, 16172C, 16223T, 16248T, 16344T, 16355T, 16519C Allentoft 2015; Mathieson 2015; *additional info from Sergey Malyshev
Afanasievo Russia Bateni [RISE510] F 2851-2468 BC 249,386



J2a2a 73G, 150T, 195C, 235G, 295T, 709A, 750G, 1438G, 3107T, 6671C, 10398G, 10499G, 11002G,11251G, 11377A, 12570G, 12612G, 13708A, 14071G, 14766T, 15257A, 15326G, 15452A, 15679G, 16069T, 16126C Allentoft 2015
Globular Amphora Poland Kowal, Kuyavia region
2850-2570 BC

K2a*
Kozlowski 2014
Yamnaya / Catacomb Russia Ulan IV, kurgan 4, grave 8 [RISE552] M 2849-2146 BC 971,022 I2a2a1b1b

[* I2a2a1b1b2]

L699; [* Malyshev reads S12195] T2a1a 73G, 263G, 709A, 750G, 1438G, 2706G, 2850C, 3107T, 4917G, 8697A, 10463C, 11251G, 11719A, 11812G, 13368A, 13965C, 14233G, 14687G, 14766T, 14905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16294T, 16296T, 16519C Allentoft 2015; Mathieson 2015; * additional info from Sergey Malyshev and Felix Immanuel
Vucedol Beli Manastir-Popova zemlja

[I3499 / GEN72]

M 2884-2666 calBCE R1b1a1a2a2

[CTS1078 / Z2103]

R1b1a1a2a2:Z2105:15747432C->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:PF6451:14116584T->A; R1b1a:A702:10038192G->A; R1b1a:FGC35:18407611C->T; R1b1a:FGC36:13822833G->T; R1b1a:FGC41:7900883C->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS5611:16394489T->G; R1:L875:16742224A->G; R1:P231:9989615A->G; R:CTS7876:17722802G->A; etc. T2e Olalde 2017
Vucedol Croatia Vucedol tell [I2792 / GEN64] M 2872-2617 calBCE 743075 G2a2a1a2a G2a2a1a2a:PF3237:17017831G->A; G2a2a1:PF3148:7816492G->A; G2a2a:PF3181:21808944C->A; G2a:CTS11463:23122426G->A; G:CTS1283:7309873T->G; G:CTS2357:14273557C->T; etc. T2c2 Olalde 2017
Bell Beaker France Hégenheim (Haut-Rhin) [I1392 / 13-Grave9] F
2832–2476 calBCE 635946 H1+152 Olalde 2017
Corded Ware Germany Bergrheinfeld [RISE446] M 2829-2465 BC 187862

R1a1a1

M417

CTS7278+, CTS10993+, FGC2547+, FGC2550+), xCTS4385 (Z2461-), xZ645 (Z647-)

U5b1c2 73G, 150T, 263G, 516T, 750G, 1438G, 2706G, 3107T, 3197C, 7028T, 9477A, 11467G, 11719A, 12308G, 12372A, 13617C, 14182C, 14766T, 15191C, 15326G, 16174T, 16189C, 16270T, 16311C Allentoft 2015; Mathieson 2015; additional info from KSDA (Anthrogenica)
Corded Ware Poland Jagodno, Wroclaw [1] M 2800 BC G?


Gworys 2013
Corded Ware Poland Jagodno, Wroclaw [2] M 2800 BC J or I?


Gworys 2013
Krotovo Russia Sopka-2/4b [Krb10]
2800-2300 BC

A10*

2 samples 663G, 16223T, 16227C, 16290T, 16311C, 16319A

Pilipenko 2015

Spain Pico Ramos, Bizkaia
2790-2100 BC

H 9 samples Izagirre and de la Rúa 1999

Spain Pico Ramos, Bizkaia
2790-2100 BC

J 4 samples Izagirre and de la Rúa 1999

Spain Pico Ramos, Bizkaia
2790-2100 BC

K 4 samples Izagirre and de la Rúa 1999

Spain Pico Ramos, Bizkaia
2790-2100 BC

U 3 samples Izagirre and de la Rúa 1999

Spain Pico Ramos, Bizkaia
2790-2100 BC

T or X 4 samples Izagirre and de la Rúa 1999
Catacomb Ukraine Krasnorechensk [KNO4]
4700-4500 y. a.

U 7028T, 11719A, 12308G, 14766T, 16356C Wilde 2014
Catacomb Ukraine Lisichansk [LIS1]
4700-4500 y. a.

U5a 7028T, 11719A, 12308G, 14766T, 16256T 16270T 16399G, reported as U Wilde 2014
Catacomb Ukraine Lisichansk [LIS2]
4700-4500 y. a.

U 7028T, 6366C, 11719A, 12308G, 13708A, 14766T, 16356C Wilde 2014
Catacomb Ukraine Lisichansk [LIS3]
4700-4500 y. a.

H 14793G, 16354T Wilde 2014
Catacomb Ukraine Shakhta Stepnaya [SAC2]
4700-4500 y. a.

J 4216C, 7028T, 10398G, 11719A, 13708A, 14766T, 16069T 16126C 16193T Wilde 2014
Catacomb Russia Temrta III [TEM2]
4700-4500 y. a.

H rCRS in HV1 Wilde 2014
Catacomb Russia Temrta V [TEM4]
4700-4500 y. a.

U/K 7028T, 11719A, 12308G, 14769G, 16256T 16270T 16399G Wilde 2014
Catacomb Russia Temrta V [TEM5]
4700-4500 y. a.

R1 4917G, 7028T, 8388C, 11719A, 14766T, 16311C Wilde 2014
Catacomb Russia Temrta V [TEM6]
4700-4500 y. a.

R1 4917G, 7028T, 8388C, 11719A, 16311C Wilde 2014
Catacomb Russia Temrta V [TEM7]
4700-4500 y. a.

U 7028T, 12308G, 14766T, 816356C Wilde 2014

Mongolia Takhilgat Uzuur-5 [TU31]
Black/dark brown hair; brown eyes
M 2742 BC Q-M242
R1b1* 16037G, 16311C, 16319A Hollard 2014
Yamnaya Ukraine Revova, kurgan 3 [R3.16] Two anthropomorphic stelae M 2731 ± 104 BC U 7028T, 12308G, 16311C Nikitin 2017
Corded Ware Germany Oberwiederstedt 2 [OBW 1]
2700-2400 BC

HV0e A16129G T16187C C16189T T16223C G16230A T16278C T16298C Brandt 2013
Corded Ware Germany Oberwiederstedt 2 [OBW 2]
2700-2400 BC

H A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Corded Ware Germany Oberwiederstedt 2 [OBW 3]
2700-2400 BC

H T16093C A16129G T16187C C16189T C16192T T16223C G16230A T16278C C16311T Brandt 2013
Corded Ware Germany Oberwiederstedt 2 [OBW 4]
2700-2400 BC

T2c T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16292T C16294T C16311T Brandt 2013
Corded Ware Germany Karsdorf [KAR 2]
2698-2658 BC

U4 A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C T16362C Brandt 2013
Corded Ware? Germany Karsdorf [KAR 24]



J1c C146T C152T G185A C195T G228A A247G C295T C16069T T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Nordic Middle Neolithic Denmark Kyndelose [RISE61] F 2650-2300 BC including reduction for high marine signal 293,152

R1a1a1

[*R1a1a1b1a3b1]

Page7+; [*Tagankin adds: CTS8401+ , Z281+] J1c4 185A, 228A, 263G, 295T, 462T, 489C, 750G, 1438G, 2706G, 3010A, 3107C, 10398G, 11251G, 11719A, 12083G, 12612G, 13708A, 14798C, 15326G, 15452A, 16069T, 16126C Allentoft 2015; Mathieson 2015; *additional info from Vladimir Tagankin
Corded Ware? Germany Karsdorf [KAR 27]



U4a1 A16129G C16134T T16187C C16189T T16223C G16230A T16278C C16311T T16356C Brandt 2013
Corded Ware Germany Eulau [EUL 26]
2628-2570 BC

H5 A16129G T16187C C16189T T16223C G16230A T16278C T16304C C16311T Brandt 2013
Battle Axe Sweden Viby [RISE94] M 2621-2472 BC 619,914

R1a1

[* R1a1a1b]

M459; Low coverage

[*KSDA reads S224/Z645+, Z649+, Z651+]

K1a2a 73G, 263G, 497T, 1438G, 1811G, 2706G 3107T, 3480G, 5773A, 9055A, 9698C, 10398G, 10550G, 11025C, 11299C, 11467G, 11719A, 12192A, 12308G, 12372A, 14167T, 14766T, 14798C, 15326G, 16189C, 16224C, 16311C, 16519C Allentoft 2015; Mathieson 2015; *add info from KSDA (Anthrogenica)
Baltic LN Lithuania Gyvakarai [1] M 2620-2470 cal BCE 1122796 R1a1a1b K1b2a Mittnik 2017
Corded Ware Germany Karsdorf [KAR 28]
2600 BC

U2e2 A16051G T16092C A16129c T16187C T16223C G16230A T16278C C16311T T16362C Brandt 2013
Corded Ware? Germany Karsdorf [KAR 30]



T2 C195T A247G T279C T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T C16296T C16311T Brandt 2013
Corded Ware Germany Karsdorf [KAR 31 and 41]
2614-2578 BC (date for sample 41)

T1a1'3 2 samples C146T A247G T16126C A16129G A16163G C16186T T16187C T16223C G16230A T16278C C16294T C16311T Brandt 2013
Corded Ware? Germany Karsdorf [KAR 38]



U5a1b1 A16129G T16187C C16189T C16192T T16223C G16230A C16256T C16270T T16278C C16291T C16311T A16399G Brandt 2013
Corded Ware Germany Karsdorf [KAR 56]
2600 BC

U5b C146T C150T C152T C195T A247G A16129G T16187C C16189T C16192T T16223C G16230A C16270T T16278C C16311T Brandt 2013
Corded Ware Germany Karsdorf [KAR 58]
2600 BC

H G73A C146T C152T C195T A247G A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Corded Ware Germany Eulau [EUL 6 (98-7)] F 2600 BC

H10e T16093C A16129G T16187C C16189T C16221T T16223C G16230A T16278C C16311T Haak 2008
Corded Ware Germany Eulau [EUL 1 (90-5)] F 2600 BC

I T16187C C16189T G16230A T16278C C16311T G16391A Haak 2008; Brandt 2013
Corded Ware Germany Eulau [EUL 3 (93-13)] ? 2600 BC

K1a24a A16129G G16145A T16187C C16189T T16223C T16224C G16230A T16278C Haak 2008; Brandt 2013
Corded Ware Germany Eulau [EUL 10 (99-1)] F 2600 BC

K1 T16093C A16129G T16187C C16189T T16223C T16224C G16230A T16278C G16319A Haak 2008; Brandt 2013
Corded Ware Germany Eulau [EUL 11 (99-2) and 12 (99-4)] M 2600 BC R1a1 SRY10831.2
Y-Search 2C46S
K1 Siblings, children of 99-1 and 99-3. T16093C A16129G T16187C C16189T T16223C T16224C G16230A T16278C G16319A
Haak 2008; Brandt 2013
Corded Ware Germany Eulau [EUL 9 (99-3)] M 2600 BC R1a1 SRY10831.2
Y-Search 2C46S
U5b1a'b A16129G T16187C C16192T T16223C G16230A C16270T T16278C C16311T Haak 2008; Brandt 2013
Corded Ware Germany Eulau [EUL 7 (98-10) and 8 (98-9)] M + F 2600 BC

X2b1'2'3'4'5'6 Siblings in same grave. C146T C152T A153G G225A T226C A247G A16129G T16187C G16230A C16311T
Haak 2008; Brandt 2013
Corded Ware? Germany Eulau [EUL 25]



T1a T16126C A16129G A16163G C16186T T16187C T16223C G16230A T16278C C16294T C16311T Brandt 2013
Corded Ware Estonia Sope [RISE00] F 4575–4350 cal BP
656226

H5a1 263G, 456T, 750G, 1438G, 3107T, 4336C, 15326G, 15833T, 16304C Allentoft 2015; Saag 2017
Bell Beaker Germany Kromsdorf [grave 3] M 2600-2500 BC

U2e 16051G, 16092C, 16129C, 16182C, 16183C, 16189C, 16362C, 73G, 152C, 263G Lee 2012
Bell Beaker Germany Kromsdorf [grave 4a] F 2600-2500 BC

W5a 16223T, 16292T, 16362C, 73G, 189G, 194T, 195C, 204C, 207A, 263G Lee 2012
Bell Beaker Germany Kromsdorf [grave 5] M 2600-2500 BC R1b1b2 M343, M269, (x S21/U106) I1a1 16129A, 16172C, 16223T, 16311C, 16391A, 73G, 199C, 203A, 204C, 250C, 263G, reported as I1. Lee 2012; Oliveiri 2013.
Bell Beaker Germany Kromsdorf [grave 8] M 2600-2500 BC R1b M343 (M269 unclear) K1 16093C, 16224C, 16311C, 16319A, 73G, 152C, 195C, 263G Lee 2012
Bell Beaker Germany Kromsdorf [grave 9a] F 2600-2500 BC

U5a1 16256T, 16270T, 16399G, 73G, 73G, 152C, 263G Lee 2012
Bell Beaker Germany Kromsdorf [grave 9b] M 2600-2500 BC

T1a 16126C, 16163G, 16186T, 16189C, 16294T, 195C, 263G Lee 2012
Corded Ware Estonia Kunila [Kunila 1] M 4800–4000 cal BP R1a-Z645 U5b1b Saag 2017
Corded Ware / Baltic LN Estonia Kunila [Kunila 2] M 2580-2340 cal BCE 382562 R1a1a1 Z283 J1c3 Mittnik 2017; Saag 2017
Yamna Ukraine Liubasha, kurgan 2, main burial L19] 2594 ± 96 BC U5 7028T, 12308G, 16270T Nikitin 2017
Sweden Ölsund, Hälsingland M 2573-2140 cal BCE 674610 R1a1a1b U4c2a Mittnik 2017
Corded Ware Germany Eulau [EUL 27]
2568-2518 BC

H2a1 A16129G T16187C C16189T T16223C G16230A T16278C C16311T C16354T Brandt 2013

Spain El Mirador, Burgos [I1277 / MIR 14] M
2568-2346 calBCE (3950±30 BP) 566,637 I2a2a
P221 H3 Mathieson 2015; Lazaridis 2016
Corded Ware Germany Esperstedt [ESP 16 / I0103] F 2566-2477 BC 1,109,474

W6a A16129G T16187C C16189T C16192T G16230A T16278C C16292T C16311T T16325C Adler 2012; Brandt 2013; Haak 2015; Lazaridis 2016
Corded Ware Germany Karsdorf [KAR 26 and 46]
2565-2521 BC (date for sample 46)

T2b4f 2 samples C146T C195T A247G T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T T16304C C16311T Brandt 2013
Corded Ware Germany Quedlinburg [QLB 23]



H C16114a A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Corded Ware Germany Esperstedt [I1534/ESP 14] M
2500-2400 BC (R1b1a2?)
(CTS11468?)
K1a1b2a A16129G T16187C C16189T T16223C T16224C G16230A T16278C Adler 2012; Brandt 2013; Mathieson 2015; but Y-DNA SNP negatively verified by Sergey Malyshev

Corded Ware

Germany Esperstedt [ESP 5 and 19]
2500-2400 BC

U5a 2 samples of A16129G T16187C C16189T C16192T T16223C G16230A C16256T C16270T T16278C C16311T Adler 2012; Brandt 2013
Corded Ware Germany Esperstedt [I1536/ESP 17] M 2500-2400 BC R1a
L62
U5a1a'g A16129G T16187C C16189T T16223C G16230A C16256T C16270T T16278C C16311T A16399G Adler 2012; Brandt 2013; Mathieson 2015
Corded Ware Switzerland Spreitenbach-Moosweg [1, 18/13] M + F 2500 BC

J1c 2 samples 73G, 185A, 228A, 263G, 295T, 315.1C, 12612. 16069T, 16126C Warnberg 2012
Corded Ware Switzerland Spreitenbach-Moosweg [3] F 2500 BC

K 73G, 195C, 263G, 315.1C, 10550, 16224C, 16311C Warnberg 2012
Corded Ware Switzerland Spreitenbach-Moosweg [2] F 2500 BC

T2b 133681, 6126C, 16294T, 16296T, 16304C Warnberg 2012
Corded Ware Switzerland Spreitenbach-Moosweg [8, 9, 15/4] M + F 2500 BC

H11a 3 samples 152C, 195C, 263G, 309.1C, 309.2C, 315.1C, 7028, 16293G, 16311C, reported as H Warnberg 2012
Corded Ware Switzerland Spreitenbach-Moosweg [11/7]
2500 BC

U2e1 11467, 16051G, 16129C, 16183C, 16189C, 16362C Warnberg 2012
Corded Ware Switzerland Spreitenbach-Moosweg [16/10] M 2500 BC

H 7028, 16189C. Warnberg 2012
Corded Ware Switzerland Spreitenbach-Moosweg [17/5] F 2500 BC

J2b1a 12612, 16069T, 16126C, 16193T, 16278T Warnberg 2012
Bell Beaker Germany Quedlinburg [QLB 19]
2500-2050 BC

U5a1a'g C146T C152T C195T A247G A16129G T16187C C16189T T16223C G16230A C16256T C16270T T16278C C16311T A16399G Brandt 2013
Corded Ware Germany Esperstedt [I1532/ESP 8] M 2500-2050 BC R1a1a L168, M512, L449 J1c2e Mathieson 2015
Corded Ware Germany Esperstedt [I1538/ESP 20] M 2500-2050 BC R1a L62 J1c5 C16069T T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Adler 2012; Brandt 2013; Mathieson 2015
Corded Ware Germany Esperstedt [I1539/ESP 25] F 2573-2511 BC J1c1b1a C16069T T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Adler 2012; Brandt 2013; Mathieson 2015
Yamna Ukraine Liubasha, kurgan 2 [L8] M 2512 ± 105 BC C4a2 16218T, 16223T, 16288C, 16298C, 16305T, 16327T, 16357C Nikitin 2017; Newton 2011
Corded Ware Germany Esperstedt [I1540/ESP 28] M 2500-2050 BC R1a1 L120 J1c5 C16069T T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Adler 2012; Brandt 2013; Mathieson 2015
Corded Ware Germany Esperstedt [I1541/ESP 32] M 2500-2050 BC R1a L62 U2e1a1 Mathieson 2015
Corded Ware Germany Esperstedt [I1542/ESP 33] M 2500-2050 BC P1 P228, P239 J2a2a Mathieson 2015
Corded Ware Germany Esperstedt [I1544/ESP 36] M 2500-2050 BC R1a L63, L146 K2b2 Mathieson 2015
Bell Beaker Germany Quedlinburg [QLB 24]
2500-2050 BC

H5 G73A C146T C152T C195T A247G A16129G T16187C C16189T T16223C G16230A T16278C T16304C C16311T Brandt 2013
Bell Beaker Germany Quedlinburg [QLB 25] 2500-2050 BC T2e T16126C A16129G G16153A T16187C C16189T T16223C G16230A T16278C C16294T C16296T C16311T Brandt 2013
Bell Beaker Germany Benzingerode-Heimburg [I1546/ BZH 2] F
2500-2050 BC

U5a1b1 A16129G T16187C C16189T C16192T T16223C G16230A C16256T C16270T T16278C C16291T C16311T A16399G Brandt 2013; Mathieson 2015
Bell Beaker Germany Benzingerode-Heimburg [I1549/BZH 15] F 2500-2050 BC

W1c1 A16129G T16187C C16189T G16230A T16278C C16292T C16311T Brandt 2013; Mathieson 2015
Corded Ware Germany Esperstedt [I0104/ESP 11] M 2473-2348 BC 990,775 R1a1a1*
M417+, M515+, M198+, L168+, M512+, M514+, L449+, CTS7083-, S466/Z280/S441-
U4b1a1a1 A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C T16362C Adler 2012; Brandt 2013; Haak 2015; Mathieson 2015
Corded Ware Germany Esperstedt [ESP 15]
2465-2395 BC

H6a1a G73A C146T C152T C195T T239C A247G A769G A825t A1018G G2706A A2758G C2885T T3594C G3915A G4104A T4312C A4727G T7028C G7146A T7256C A7521G T8468C T8655C G8701A G9380A C9540T G10398A T10664C A10688G C10810T C10873T C10915T T11253C A11719G A11914G T12705C G13105A G13276A T13506C T13650C T14766C A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16362C A16482G Brotherton 2013; Brandt 2013
Corded Ware Germany Esperstedt [ESP 22 / I0049] F 2464-2210 calBCE (3867±35 BP) 534,318

X2b4 A16129G T16187C G16230A C16311T Adler 2012; Brandt 2013; Haak 2015; Lazaridis 2016
Corded Ware Germany Esperstedt [ESP 26 / I0106] F 2464-2210 calBCE (3867±32 BP) 230,482

T2a1b1 T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T C16311T T16324C Adler 2012; Brandt 2013; Haak 2015; Lazaridis 2016
Catacomb Ukraine Nevskoe [NEV1]
4500-4000 y ago

U 7028T, 11719A, 12308G, 14766T, 14793G, 16192T 16256T 16270T 16399G Wilde 2014
Catacomb Ukraine Nevskoe [NEV3]
4500-4000 y ago

H 16362C Wilde 2014
Catacomb Ukraine Novozvanovka II [NOZ1, NOZ2]
4500-4000 y ago

U 2 samples of 7028T, 11719A, 12308G, 14766T, 16356C Wilde 2014
Catacomb Russia Peschanyi [PEJ2]
4500-4000 y ago

H 16234T Wilde 2014
Catacomb Russia Peschanyi [PEJ3]
4500-4000 y ago

H 16362C Wilde 2014
Catacomb Russia Peschanyi [PEJ4]
4500-4000 y ago

H 16362C Wilde 2014
Catacomb Russia Peschanyi [PEJ5]
4500-4000 y ago

U 7028T, 11719A, 12308G, 14766T, 16356C Wilde 2014
Catacomb Ukraine Kirovograd Sugokleya [SUG5]
4500-4000 y ago

H 16362C Wilde 2014
Catacomb Russia Temrta III [TEM1]
4500-4000 y ago

U 7028T, 11719A, 12308R, 16356C Wilde 2014
Catacomb Russia Temrta III [TEM3]
4500-4000 y ago

J1 462T, 3010A, 4216C, 7028T, 8269A, 10398G, 11719A, 13708A, 14766T, 16069T 16126C 16145A 16172C 16222T 16261T Wilde 2014
Catacomb Russia Temrta V [TEM8]
4500-4000 y ago

U 7028T, 11719A, 12308G, 14766T, 14793G Wilde 2014
Catacomb Moldova Tetcani [TET1]
4500-4000 y ago

I 4529T, 7028T, 8251A, 10398G, 11719A, 12705T, 14766T, 216223T 16391A Wilde 2014
Catacomb Ukraine Vinogradnoe [VIN3]
4500-4000 y ago

U 11719A, 12308G, 14793G, 16256T 16270T 16399G Wilde 2014
Catacomb Ukraine Vinogradnoe [VIN8]
4500-4000 y ago

J1 462T, 3010A, 4216C, 7028T, 8269A, 10398G, 11719A, 13708A, 14766T, 16069T 16126C 16145A 16172C 16222T 16261T Wilde 2014
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I0263] F 2850–2250 (2474-2300) BCE 31569 X2b+226 Olalde 2017
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I1553] F 2850–2250 (2474-2300) BCE 37205 pre-H103 Olalde 2017
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I0825] M 2850–2250 (2474-2300) BCE 42018 G2 G2:F1189:8427005A->G; G:CTS9894:19124322A->T; G:L522:17533325A->C; G:M3470:7830068T->C; G:M3487:8602816G->C K1a4a1 Olalde 2017
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I0260] F 2850–2250 (2474-2300) BCE 51816 K1a2a Olalde 2017
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I0261] M 2850–2250 (2474-2300) BCE 52264 R1b1a (xR1b1a1a2a) R1b1a:L1345:21558298G->T; R:F652:23631629C->A; R:M651:9889199G->A U5b1i Olalde 2017
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I0823] F 2850–2250 (2474-2300) BCE 60463 H1 Olalde 2017
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I0258] F 2850–2250 (2474-2300) BCE 62877 H1q Olalde 2017
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I0262] F 2850–2250 (2474-2300) BCE 92174 U5b3 Olalde 2017
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I0826] M 2833–2480 calBCE 112957 I2a2 I2a2:P218:17493630T->G; I:CTS11540:23156725C->T; I:PF3817:21939618G->A H1t Olalde 2017
Battle Axe Sweden Linkoping [Ber2]
2800–2100 BC

N1a1a1a 16147A 16172C 16223 T 16248T 16320T 16355T Malmstrom 2015
Bell Beaker Spain Arroyal, Burgos [I0459 / Roy2 / UE25] F
2600–2200 BCE 527866 U5b2b Olalde 2017
Bell Beaker Germany Osterhofen-Altenmarkt [RISE563 / I4144] M 2572–2512 calBCE


271735

R1b1a1a2a1a2

[P312 / PF6547 / S116]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:PF6463:16183412C->A; R1b1a:FGC36:13822833G->T; R1b1a:L754:22889018G->A; R1:CTS3123:14674176A->C; etc K1c1 73G, 146C, 152C, 263G, 750G, 1438G, 1811G, 2706G, 3107T, 9055A, 9698, 10398G, 10550G, 11299C, 11377A, 11467G, 11719A, 12308G, 12372A, 13708A, 14167T, 14766T, 14798C, 15326G, 16224C, 16311C, 16519C Allentoft 2015; Mathieson 2015; Olalde 2017
Bell Beaker Spain Paris Street, Cerdanyola, Barcelona [I0257] M 2571–2350 calBCE 107954 R1b1 R1b1:L1349:22722580T->C; R:L1347:22818334C->T; R:P224:17285993C->T H1ax Olalde 2017
Bell Beaker Poland Samborzec, Małopolska [I4253 / RISE1124 / grave no. 13] M 2571-2208 BCE 452974 R1b1a1a2

[M269 / PF6517]

R1b1a1a2:CTS8728:18167403C->T; R1b1a1a2:PF6399:2668456C->T; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a:A702:10038192G->A; R1b1a:FGC41:7900883C->A; R1b1a:PF6263:21159055C->A; R1b1:CTS2134:14193384G->A; R1b1:L780:21183643A->G; R1b:M343:2887824C->A; R1:CTS2565:14366723C->T; etc U5a2c Olalde 2017
Bell Beaker France Sierentz - Les Villas d'Aurčle (Haut-Rhin) [I1390 / 11-Grave68] M 2566–2299 calBCE 449568 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS8665:18137831T->C; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6505:21784286G->A; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6524:23452965T->C; R1b1a:CTS8436:18026855G->A; R1b1a:FGC35:18407611C->T; R1b1a:L754:22889018G->A; R1b1a:L1345:21558298G->T; R1b1:CTS2134:14193384G->A; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; etc X2b4a Olalde 2017
Bell Beaker Hungary Budakalász, Csajerszke [I3528 / GEN85 / Grave276] M
2559–2301 calBCE 224817 G2a2a1a2a1 G2a2a1a2a1:Z6277:13814902C->T; G2a2a1a2a:PF3238:17256431G->A; G2a:F2529:17571517A->G; G2a:F3088:20813445G->A; G2a:PF3141:23973594T->G; G2:CTS1900:14116322T->A; G2:CTS6316:16817402C->T; G2:F1239:8482393C->T; G2:L89:7978725C->T; G:CTS34:2681740G->A; etc HV4a1+16291 Olalde 2017

Portugal Sao Paulo [SP 18861]
2500 BC

H CRS, -7025, -11719, -12308 Afonso 2010

Portugal Sao Paulo [SP 9222*]
2500 BC

H, HV* or R0 CRS, -12308 Afonso 2010

Portugal Sao Paulo [SP 5730, 8001, 18248]
2500 BC

H 3 samples CRS, -7025. Afonso 2010
Bell Beaker Hungary Budakalász, Csajerszke [I3529 / GEN86 / Grave597] F
2500-2200 BCE 560151 I3a Olalde 2017
Bell Beaker Hungary Szigetszentmiklós, Felső Ürge-hegyi dűlő [I4178 / GEN_58, Grave 552] M
2500-2200 BCE 587361 R1b1a1a2

[M269]

R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS8728:18167403C->T; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6497:21222868C->G; R1b1a1a2:PF6500:21410840G->T; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6524:23452965T->C; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:FGC36:13822833G->T; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L278:18914441C->T; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS2565:14366723C->T; etc

J1c1b1a Olalde 2017
Bell Beaker Germany Irlbach, County of Straubing-Bogen, Bavaria [I4248 / RISE916, grave 3] M
2500-2000 BCE 5722 no data T2b+152 Olalde 2017
Bell Beaker Germany Irlbach, County of Straubing-Bogen, Bavaria [I4249 / RISE917, grave 4] F
2500-2000 BCE 32814 H5a1 Olalde 2017
Bell Beaker Germany Irlbach, County of Straubing-Bogen, Bavaria [I4250 / RISE918, grave 5] F
2500-2000 BCE 29790 U5a1a2b Olalde 2017
Bell Beaker Germany Manching-Oberstimm, Bavaria [I4123 / RISE556 / F0003 / Grave 4] F
2500-2000 BCE 100951 V Olalde 2017
Bell Beaker
Germany

Rothenschirmbach [ROT 6/I0108]

2497-2436 BC

H5a3 G73A C146T C152T C195T A247G C456T G513A A769G A825t A1018G G2706A A2758G C2885T T3594C G4104A T4312C T4336C T7028C G7146A T7256C A7521G T8468C T8655C G8701A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C, G13105A G13276A T13506C T13650C T14766C G15884A A16129G T16187C C16189T T16223C G16230A T16278C T16304C C16311T C16519T Adler 2012; Brotherton 2013; Brandt 2013; Haak 2015; Mathieson 2015
Bell Beaker Germany Alberstedt [ALB 2]
2494-2344 BC

H G73A C146T C152T C195T A247G A16129G T16187C C16189T T16223C G16230A T16278C C16311T Adler 2012; Brandt 2013
Bell Beaker England Amesbury (Boscombe) Down, Wiltshire [ I2457 / 13382] M
2480–2031 calBCE 718630 R1b1a1a2a1a2c

[L21]

R1b1a1a2a1a2c:L21:15654428C->G; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L52:14641193C->T; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; I4a Olalde 2017
Bell Beaker France La Fare (Forcalquier) [I2575 / Grave S. 14 ] M 2476–2211 calBCE 27777 no data K1c1 Olalde 2017
Corded Ware / Bell Beaker mix? Germany Alberstedt [I0118/ALB 3] F 2471-2246 BC
HV6'17 Haak 2015; Mathieson 2015
Bell Beaker England Amesbury (Boscombe) Down, Wiltshire [I2565 / 1238] Companion M
2470–2140 calBCE 14794 R1b1a1a2a1a2c

[L21]

R1b1a1a2a1a2c:L21:15654428C->G; R1b1a:L761:16773870A->G W1+119 Olalde 2017
Bell Beaker Hungary Budapest-Békásmegyer, Királyok útja [I2364 / GEN_10a, Grave 219/B] M
2470–2060 calBCE 767419 H2 H2:L279:6932824G->T; H2:L285:21869856C->T; H2:P96:14869743C->A; H:M2896:16919982C->T; etc U5a2b Olalde 2017
Bell Beaker Hungary Budapest-Békásmegyer, Királyok útja [I2365 / GEN_11a, Grave452] M
2465-2205 calBCE 787155 R1b1a1a2a1a2b1

[U152 > L2]

R1b1a1a2a1a2b1:L2:5755550C->T; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L11:17844018T->C; R1b1a1a2a1a:L52:14641193C->T; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS8728:18167403C->T; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:L1353:19179540G->A; R1b1a1a2:PF6409:4352151G->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6463:16183412C->A; R1b1a1a:PF6524:23452965T->C; R1b1a1:L388:17400785G->A; R1b1a:A702:10038192G->A; R1b1a:FGC35:18407611C->T; R1b1a:L761:16773870A->G; R1b1a:PF6271:23984056G->A; R1b1:CTS2229:14226692T->A; R1b1:L278:18914441C->T; R1b1:L1349:22722580T->C; R1b1:M415:9170545C->A; R1b:M343:2887824C->A; R1:CTS3123:14674176A->C; etc V3 Olalde 2017
Bell Beaker France Sierentz - Les Villas d'Aurčle (Haut-Rhin) [I1389 / 10-Grave69] M 2468–2278 calBCE 97913 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2:PF6425:7762947T->C; R1:CTS2565:14366723C->T; R:CTS207:2810583A->G; X2b4a Olalde 2017
Bell Beaker Germany Quedlinburg [I0805 / QLB 26] M
2467-2142 BC 231,622 R1b1a2
PF6430, PF6482, PF6500, PF6509
H1 Mathieson 2015; Lazaridis 2016
Bell Beaker Spain Arroyal, Burgos [I0462 / Roy5 / SU25] F
2465–2211 calBCE 32916 K1a+195 Olalde 2017
Bell Beaker England Hasting Hill (Sunderland, Tyne and Wear) [I2612 / TWCMS Sk2008-1953, find 9] F
2465–2209 calBCE 55173 U5b2b2 Olalde 2017
Bell Beaker Poland Samborzec, Małopolska [I4252 / RISE1123 / grave no. 1] M 2463-2142 BCE 30131 R R:F459:18017528G->T U5a1a1

Olalde 2017
Bell Beaker Spain Arroyal, Burgos [I0460 / Roy3 / SU25] M
2461–2210 calBCE 344361 I2a2a I2a2a:L36:17570599C->T; I2a2a:P219:15517851T->G; I2a2:L181:19077754G->T; I2a2:M436:18747493G->C; I2a:L460:7879415A->C; I:CTS1800:14073053G->A; I:CTS2536:14352669G->A; I:CTS4088:15389836T->C;

H45 Olalde 2017
Bell Beaker Germany Hugo-Eckener-Straße, Augsburg [E09568_d / Grave 8 Skeleton 1] M 2461–2210 calBCE 32968 R1b1a1a2a1a [L151] R1b1a1a2a1a:L52:14641193C->T; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS8728:18167403C->T; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:F1794:14522828G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L500:18180446C->A; R1b1a1a2:L773:7220727A->G; R1b1a1a2:M269:22739367T->C; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6505:21784286G->A; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:P297:18656508G->C; R1b1a1a:PF6463:16183412C->A; R1b1a1a:PF6498:21312064C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1:CTS2229:14226692T->A; R1b1:L278:18914441C->T; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS2565:14366723C->T; etc U5b2b Olalde 2017
Bell Beaker Germany Hugo-Eckener-Straße, Augsburg [E09613_d / Grave 3] F 2289–2141 calBCE 174567 H1 Olalde 2017
Bell Beaker Germany Hugo-Eckener-Straße, Augsburg [E09614_d / Grave 10] F 2268–2046 calBCE 46955 J1c3 Olalde 2017
Bell Beaker Germany Unterer Talweg 58-62, Augsburg [E09537_d / Feature 68 Skeleton 2] F 2464–2212 calBCE 87970 K1a Olalde 2017
Bell Beaker Germany Quedlinburg [QLB 38]
2460-2280 BC

H A16129G T16187C T16223C G16230A T16278C C16311T Brandt 2013
Bell Beaker England Amesbury (Boscombe) Down, Wiltshire [I2416 / 25004] Boscombe Bowmen M
2460-2200 BC 136956 R1b1a1a2a1a

[L151]

R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a:PF6459:15286480G->C; R1b1a:FGC36:13822833G->T; R1b1a:PF6263:21159055C->A; R1b1:CTS2229:14226692T->A; R1b:M343:2887824C->A; R1:P286:17716251C->T; R:CTS8311:17930099C->A; R:FGC1168:15667208G->C; R:P280:21843090C->G
K1b1a1 Olalde 2017
Bell Beaker England Abingdon Spring Road cemetery, Oxfordshire [I2450 / SK 3036 (ABSRC00)] F 2460–2200 calBCE 734749 K1a1b1g Olalde 2017
Bell Beaker England Yarnton, Oxfordshire [I2446 / SK 8784 (YCF 95)] F 2460-2140 calBCE 83832 K1b1a1 Olalde 2017
Spain Camino del Molino, Caravaca de la Cruz, Murcia [I0453 / Cmol79] M 2460–2140 calBCE 42882 I I:CTS4848:15862842C->T; I:CTS6497:16939794A->T; I:PF3666:8728974T->G V Olalde 2017
Bell Beaker England Amesbury (Boscombe) Down, Wiltshire [I2459 / 85684_62014] F
2460–2140 calBCE 519046 T2a1a Olalde 2017
Bell Beaker France Dolmen of Villard (Lauzet-Ubaye) [I3874 / Vil-Lauz-1435] F 2459–2242 BCE 652965 T2b3 Olalde 2017
Bell Beaker France Dolmen of Villard (Lauzet-Ubaye) [I3875 / Vil-Lauz-1316] M 2459–2242 BCE 792712 R1b1a1a2a1a2b1

[U152 > L2]

R1b1a1a2a1a2b1:L2:5755550C->T; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:M520:4446430T->A; R1b1a1a2:PF6399:2668456C->T; R1b1a1a2:PF6425:7762947T->C; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6497:21222868C->G; R1b1a1a2:PF6505:21784286G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS7941:17755905G->A; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6459:15286480G->C; R1b1a1a:PF6463:16183412C->A; R1b1a1:L388:17400785G->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:CTS8436:18026855G->A; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:L1068:21528257T->C; R1b1a:L1345:21558298G->T; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; etc Olalde 2017
Bell Beaker Hungary Szigetszentmiklós, Felső Ürge-hegyi dűlő [I2786 / GEN_56, Grave 133] M
2459–2206 calBCE 936369 I2a2a I2a2a:M223:21717307G->A; I2a2:L37:17516123T->C; I2a2:L181:19077754G->T; I2a2:L368:6931594C->T; I2a2:P216:13992338C->G; I2a:L460:7879415A->C; I2:L68:18700150C->T; I2:PF3664:8567995G->A; I:CTS48:2688442T->A; etc

I1 Olalde 2017
Bell Beaker Spain Arroyal, Burgos [I0458 / Roy1 / SU25] M
2458–2206 calBCE 131125 I2a2a I2a2a:P220:24475669G->T; I2a2:M436:18747493G->C; I2:PF3664:8567995G->A; I:CTS6497:16939794A->T; I:CTS8333:17940414G->A; I:CTS8742:18172947A->G; I:FGC2412:21689728A->G; I:PF3836:22525421T->G K1a1b1 Olalde 2017
Bell Beaker Hungary Szigetszentmiklós, Felső Ürge-hegyi dűlő [I2741 / GEN_20, Grave 49] M
2458–2154 calBCE 871912 I2a1a1 I2a1a1:L158:23496560G->A; I2a1a1:M26:21865821G->A; I2:L68:18700150C->T; I2:PF3664:8567995G->A; I:CTS48:2688442T->A; etc H1+16189 Olalde 2017
Bell Beaker Hungary Szigetszentmiklós, Felső Ürge-hegyi dűlő [I2787 / GEN_59, Grave 688] M
2458–2202 calBCE 669314 R1b1a1a2a2

[CTS1078 / Z2103]

R1b1a1a2a2:CTS1078:7186135G->C; R1b1a1a2a2:Z2105:15747432C->A; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS8728:18167403C->T; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L1353:19179540G->A; R1b1a1a2:PF6399:2668456C->T; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6500:21410840G->T; R1b1a1a2:PF6505:21784286G->A; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:L752:18394634T->C; R1b1a1a:PF6451:14116584T->A; R1b1a1a:PF6459:15286480G->C; R1b1a1a:PF6463:16183412C->A; R1b1a1a:PF6498:21312064C->A; R1b1a:CTS3063:14637352T->C; R1b1a:CTS8436:18026855G->A; R1b1a:FGC35:18407611C->T; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1a:L1068:21528257T->C; R1b1a:L1345:21558298G->T; R1b1a:PF6249:8214827C->T; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS3625:15080010C->T; R1b1:L278:18914441C->T; R1b1:M415:9170545C->A; R1b:M343:2887824C->A; R1:CTS2565:14366723C->T; etc T2b Olalde 2017
Bell Beaker? Germany Quedlinburg [QUEXII 3 in Brandt, I0112 / QUEXII 6 in Haak] F 2457-2142 calBCE (3820±42 BP) 985,107

H13a1a2c G73A C146T C152T C195T A247G A769G A825t A1018G C2259T G2706A A2758G C2885T T3594C G4104A T4312C A4745G T7028C G7146A T7256C A7521G T8468C T8655C G8701A G9025A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C A13542G T13650C C13680T T14766C C14872T A16129G T16187C C16189T T16223C G16230A T16278C C16311T C16519T Adler 2012; Brotherton 2013; Brandt 2013; Haak 2015; Lazaridis 2016
Bell Beaker France Marlens - Sur les Barmes (Haute-Savoie) [I1388 / 9-Grave1] M
2456–2135 calBCE 375748 R1b1a1a2a1a

[L151]

R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:PF6463:16183412C->A; R1b1a1a:PF6498:21312064C->A; R1b1a:A702:10038192G->A; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A;

H Olalde 2017
Chalcolithic Portugal Galeria da Cisterna (Almonda karst system), Torres Novas, [I0840 / AMD2-G18-187] F 2456–2201 calBCE 133290 U5b1i Olalde 2017
Bell Beaker England Amesbury (Boscombe) Down, Wiltshire [I2418 / 6033 ] F
2440–2200 calBCE 656573 K1a4a1 Olalde 2017
Catacomb Ukraine Revova, kurgan 3 [R3.13] 2437 ± 95 BC U5 7028T, 12308G, 16270T Nikitin 2017
Catacomb Ukraine Dubinovo, kurgan 1 [D1.8] M 2435 ± 106 BC U5a1a1h 7028T,12308G, 16192T, 16256T, 16270T, 16294T, 16296T; previously reported as C4a3 by Newton 2011 Nikitin 2017;Newton 2011
Bell Beaker France Mondelange - PAC de la Sente (Moselle) [I1382 / 3-Grave515] M 2435–2136 calBCE 607837 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:PF6459:15286480G->C; R1b1a1a:PF6463:16183412C->A; R1b1a:CTS4244:15510064T->G; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS997:7132713G->A;

U5a2c3a Olalde 2017
Bell Beaker Germany Quedlinburg [I0806 / QLB 28] Frequently rode horses. M 2431-2150 calBCE (3824±25 BP) 142,638 R1b1a1a2a1a2

[P312]

P312+, S1217-, Z262-, DF49-, DF23-, L554-, S868-, CTS6581-, CTS2457.2- [* ZZ11+ equivalent (ancestral to DF27 and U152); DF27+ ? in 390k BAM file, but short and may actually belong to chromosomes 2 or 5.] H1 G73A C146T C152T C195T A247G A769G A825t A1018G G2706A A2758G C2885T G3010A T3594C G4104A T4312C T7028C G7146A T7256C A7521G T8468C T8655C G8701A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C T14766C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brotherton 2013; Brandt 2013; Haak 2015; Mathieson 2015; Lazaridis 2016;* additional information from Alex Williamson
Bell Beaker Germany Rothenschirmbach [ROT 3/I0060] F 2428-2149 BC K1a2c Mathieson 2015
Bell Beaker Germany Rothenschirmbach [ROT 4/I0111] F 2414-2333 BC

H3a0 A16129G T16187C C16189T T16223C G16230A C16256T T16278C C16311T Brandt 2013; Haak 2015; Mathieson 2015
Bell Beaker Poland Samborzec, Małopolska [I4251 / RISE1122/ grave no. 7] 2400-2200 BC* 80714 R1b1a1a2

[M269]

R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:L150.1:10008791C->T; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1:P238:7771131G->A H1 Olalde 2017; *personal communication from Piotr Włodarczak, correcting older radiocarbon date given in Olalde as from an unreliable lab
Bell Beaker England Yarnton, Oxfordshire [I2447 / SK 8779 (YCF 95)] M 2400–2040 calBCE 702581 R1b1a1a2a1a2c1

[L21 > DF13]

R1b1a1a2a1a2c1:CTS8221:17885577C->T; R1b1a1a2a1a2c:L21:15654428C->G; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8728:18167403C->T; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:L752:18394634T->C; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1a:L1345:21558298G->T; R1b1a:PF6263:21159055C->A; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L278:18914441C->T; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; etc Olalde 2017
Bell Beaker France Mondelange - PAC de la Sente (Moselle) [I1381 / 2-Grave487] M 2400–1900 BCE 690979 R1b1a1a2a1a

[L151]

R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:L752:18394634T->C; R1b1a:A702:10038192G->A; R1b1a:FGC35:18407611C->T; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L780:21183643A->G; R1b1:L822:7960019G->A; R1b1:M415:9170545C->A; R1b:M343:2887824C->A; R1:CTS997:7132713G->A;

H Olalde 2017
Bell Beaker England Staxton Beacon (North Yorkshire) [I1770 / Burial11] F
2400–1600 calBCE 483474 U5b1d2 Olalde 2017
Yamna Ukraine Revova, kurgan 3 [R3.7] 2389 ± 81 BC U5a 7028T, 12308G, 16192T, 16256T,16270T Nikitin 2017
Bell Beaker Germany Alberstedt [ALB 1]
2371-2282 BC

H3b G73A C146T C152T C195T A247G A769G A825t A1018G A2581G G2706A A2758G C2885T T3594C G4104A T4312C T6776C T7028C G7146A T7256C A7521G T8468C T8655C G8701A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C T14766C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brotherton 2013; Brandt 2013
Catacomb Ukraine Dubinovo, kurgan 1 [D1.11] F 2369 ± 111 BC 7028T, 12308G Nikitin 2017
Catacomb Ukraine Dubinovo, kurgan 1 [D1.10] n.d. U5 7028T, 12308G, 16270T Nikitin 2017
Bell Beaker Germany Eulau [EUL 23]
2351-2282 BC

H G73A C146T C152T C195T A247G G263A A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Bell Beaker Germany Ausburg [RISE559 / I4124] F 254589 H46 152C, 263G, 1438G, 2478A, 2772T, 9281T, 11875C, 15326G, 16192T, 16519C Allentoft 2015; Olalde 2017 increased coverage
Bell Beaker Germany Ausburg [RISE560 / I4132] M 223736 R1b1a1a2a1a [L151 / PF6542, L52 / PF6541, P310] R1b1a1a2a1a:L52:14641193C->T; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a2:PF6505:21784286G->A; R1b1a1a2:PF6509:22190371A->G; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6463:16183412C->A; R1b1a:CTS4244:15510064T->G; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; R:CTS3622:15078469C->G; etc U5a1a1 73G, 263G, 750G, 886T, 1700C, 2706G, 3197C, 5495C, 11467G, 12308G, 12372A, 13617C, 14766T, 14793G, 15218G, 15326G, 15924G, 16256T, 16270T, 16311C, 16399G Allentoft 2015; Mathieson 2015; Olalde 2017 increased coverage and resolution Y-DNA
Bell Beaker Germany Landau an der Isar [RISE562] F
68,118



H2a1e 263G, 575T, 652A, 951A, 1845T, 5326T, 15326G, 16354T Allentoft 2015
Bell Beaker Germany Osterhofen-Altenmarkt [RISE564] M
93,003

R1b1a1a2a1 L51/M412/PF6536/S167+ (no reads on L11, P312, U152) H+T16311C 263G, 750G, 1438G, 2706G, 3107T, 4770A, 7028T, 14605G, 15326G, 16038G, 16213A, 16311C Allentoft 2015; Mathieson 2015; additional info from Richard Rocca and Felix Emmanuel.
Bell Beaker Spain Arroyal, Burgos [I0461 / Roy4 / SU19] F
2348–2200 calBCE 720612 K1a1b1 Olalde 2017
Bell Beaker France Rouffach – Rue de Pfaffenheim (Haut-Rhin) [I1391 / 12-Grave Excavataion2014] F
2346–2133 calBCE 87166 J1c4 Olalde 2017
Bell Beaker Germany Rothenschirmbach [ROT 1/ I1530] M
2345-2198 BC R1b1a1a2 P233 [CTS2134; CTS623] H3ao2 G73A C146T C152T C195T A247G A769G A825t A1018G G2706A A2758G C2885T T3594C G4104A T4312C C4577T T6776C T7028C G7146A T7256C A7521G T8468C T8655C G8701A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C T14766C A16129G T16187C C16189T T16223C G16230A C16256T T16278C C16311T Adler 2012; Brandt 2013; Mathieson 2015; [ additional SNPs from Richard Rocca]
Bell Beaker? Germany Benzingerode-Heimburg [BZH 6 / I0059] F 2337-2138 calBCE (3796±30 BP, MAMS-21486) 752,357

H1 / H1b'ad G73A C146T C152T C195T A247G A769G A825t A1018G G2706A A2758G C2885T G3010A T3594C G4104A T4312C T7028C G7146A T7256C A7521G A8149G T8468C T8655C G8701A A9377G T9467C C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C A13671G T14319C T14766C A16129G T16187C T16223C G16230A T16278C C16311T Brotherton 2013; Brandt 2013; Haak 2015; Lazaridis 2016
Bell Beaker England Windmill Fields, Ingleby Barwick (Stockton-on-Tees, County Durham) [I1765_d/Sk7] F 2330–2040 calBCE 13484 N1a1a1a2 Olalde 2017
Bell Beaker Germany Karsdorf [KAR 51]
2314-2278 BC

U4c1 C146T C152T A247G A16129G C16179T T16187C C16189T T16223C G16230A T16278C C16311T T16356C Brandt 2013
Bell Beaker Germany Bruck, City of Künzing, County of Deggendorf, Bavaria [I3604 / Obj. 278, grave 9] Metalworker M
2300 BCE 30589 R1b1a R1b1a:A702:10038192G->A; R:L1225:22733758C->G; R:M613:7133986G->C HV9 Olalde 2017
Bell Beaker Germany Bruck, City of Künzing, County of Deggendorf, Bavaria [I3607 / Obj. 335, grave 7] M
2300 BCE 10064 R1b1a1a

[P297 / PF6398]

R1b1a1a:CTS5577:16376495A->C; R1b1a:L754:22889018G->A W3a1 Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3601 / Grave 15] F
2300–2150 BCE 656910 H10e Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3602 / Grave 16] F
2300–2150 BCE 85941

I3a

Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3600 / Grave 14] F
2300–2150 BCE 288583 U5b3

Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3599 / Grave 13] Ist-degree relative of I3588 M
2300–2150 BCE 36524 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1:CTS5611:16394489T->G H1e1a Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3588 / Grave 2] Ist-degree relative of I3599 M
2300–2150 BCE 428761 R1b1a1a2a1a

[L151 / PF6542]

R1b1a1a2a1a:P310:18907236A->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:PF6459:15286480G->C; R1b1a:A702:10038192G->A; R1b1a:FGC41:7900883C->A; R1b1a:L1345:21558298G->T; R1b:M343:2887824C->A; R1:CTS2565:14366723C->T; etc

H1e1a Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3589 / Grave 3] M
2300–2150 BCE 547552 R1b1a1a2a1a2b1

[U152 > L2]

R1b1a1a2a1a2b1:L2:5755550C->T; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a:CTS4244:15510064T->G; R1b1a:L761:16773870A->G; R1b1a:L1345:21558298G->T; R1b1a:PF6249:8214827C->T; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L780:21183643A->G; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; R1:CTS2565:14366723C->T; etc U4d1 Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3590 / Grave 4] F
2300–2150 BCE 437017 H1 Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3594 / Grave 9]
2300–2150 BCE 636256 T2f Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3592 / Grave 8] F
2300–2150 BCE 18087 H+16129 Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3593 / Grave 6]
2300–2150 BCE 80945 H1 Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3597 / Grave 12] M
2300–2150 BCE 121613 R1b1a1a2a1a2b1

[U152 > L2]

R1b1a1a2a1a2b1:L2:5755550C->T; R1b1a1a2:PF6509:22190371A->G; R1b1a1a:FGC57:7759944G->A; R:CTS8311:17930099C->A; etc

T2f Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3587 / Grave 1] U
2300–2150 BCE 8020 H1e1a Olalde 2017
Bell Beaker Germany Alburg, Lerchenhaid-Spedition Häring, City of Straubing, Bavaria [I3596 / Grave 11] M
2300–2150 BCE 7616 no data H+16129 Olalde 2017
Corded Ware Germany Quedlinburg [QUEXII 1] F 2300-2130 BC

H4a1 G73A C146T C152T C195T A247G A769G A825t A1018G G2706A A2758G C2885T T3594C C3992T A4024G G4104A T4312C T5004C T7028C G7146A T7256C A7521G T8468C T8655C G8701A G9123A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C C14365T A14582G T14766C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Adler 2012; Brandt 2013
Odinovo Russia Preobrajenka- 6 [Od7 and Od11]
2300-2000 BC

A10*

2 samples 663G, 16223T, 16227C, 16290T, 16311C, 16319A

Pilipenko 2015
Odinovo Russia Sopka-2/4a [Krz1]
2300-2100 BC

A10a 663G, 16223T, 16227C, 16230G, 16290T, 16311C, 16319A Pilipenko 2015
Bell Beaker Germany Quedlinburg [QLB 27]
2300-2130 BC

U5b A16129G T16187C G16213A T16223C G16230A C16270T T16278C C16311T Brandt 2013
Bell Beaker Netherlands Oostwoud-Tuithoorn, West Frisia [I4075 / skeleton 243-M15] F 2300–1900 BCE 770037 H5a1 Olalde 2017
Bell Beaker Germany Rothenschirmbach [ROT 3] F 2294-2206 BC

K1a2c T16086C A16129G T16187C C16189T T16223C T16224C G16230A T16278C Brandt 2013; Haak 2015
Bell Beaker Germany Quedlinburg [QUEXII 4] F 2290-2130 BC

J1c5 C16069T T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Adler 2012; Brandt 2013; Haak 2015
Bell Beaker England Yarnton, Oxfordshire [I2443 / SK 8772 (YCF 95), 2] F 2290–1980 calBCE 806658 T2c1d+152 Olalde 2017
Chalcolithic Portugal Cova de Moura,Torres Vedras [I4229 / CDM3] M 2289–2135 calBCE 810327 I2a1a1 I2a2a:P220:24475669G->T; I2a2:M436:18747493G->C; I2:PF3664:8567995G->A; I:CTS6497:16939794A->T U5a2+16362 Olalde 2017
Bell Beaker England West Deeping, Lincolnshire [I2453/CQWDO7, feature F.320 skeleton 1126] M 2289–2041 calBCE 610725 R1b1a1a2a1a2c1

[L21 > DF13]

R1b1a1a2a1a2c1:CTS8221:17885577C->T; R1b1a1a2a1a::14641193C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6497:21222868C->G; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6498:21312064C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1:CTS2134:14193384G->A; R1b1:M415:9170545C->A; R1b:M343:2887824C->A; etc K2a Olalde 2017
Scotland Dryburn Bridge, East Lothian [I2568 / GENSCOT15, Burial 10, Cist 2] M 2287–2039 calBCE 570382 R1b1a1a2a1a2c1

[L21 > DF13]

R1b1a1a2a1a2c1:CTS8221:17885577C->T; R1b1a1a2a1a2c:L21:15654428C->G; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a2:PF6509:22190371A->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:L1345:21558298G->T; R1b1a:PF6249:8214827C->T; R1b1a:PF6263:21159055C->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L278:18914441C->T; R1b1:L780:21183643A->G; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; etc. U5a1b1g Olalde 2017
Bell Beaker? Germany Benzingerode-Heimburg [BZH 4] F 2283-2146 BC

H1e G73A C146T C152T C195T A247G A769G A825t A1018G G2706A A2758G C2885T G3010A T3594C G4104A T4312C G5460A T7028C G7146A T7256C A7521G T8468C T8655C G8701A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C T14766C A15220G A15401G A16129G T16187C C16189T T16223C G16230A T16278C A16293G C16311T Brotherton 2013; Brandt 2013; Haak 2015
Chalcolithic Portugal Cova de Moura,Torres Vedras [CM364] M 36493410 I2a1b H Martiniano 2017
Chalcolithic Portugal Cova de Moura,Torres Vedras [CM9B] F 113150709 J1c1 Martiniano 2017
Bell Beaker Spain Camino de las Yeseras (San Fernando de Henares, Madrid) [I4245 / RISE695] F
2280–1790 BCE 14942 L1b1a Olalde 2017
Bell Beaker Spain Camino de las Yeseras (San Fernando de Henares, Madrid) [I4247 / RISE698] F
2280–1790 BCE 188194 T2b3 Olalde 2017
BB? England Amesbury (Boscombe) Down, Wiltshire [I2596 / 5289] M
2280–2030 calBCE 354928 R1b1a1a2a1a

[L151 / PF6542 / P311]

R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a:CTS4244:15510064T->G; R1b1:CTS2134:14193384G->A; R1:L875:16742224A->G; R1:M306:22750583C->A; R:CTS8311:17930099C->A; R:F295:15594523A->G; R:F459:18017528G->T; R:FGC1168:15667208G->C; R:L1225:22733758C->G; R:L1347:22818334C->T J1c1 Olalde 2017
Bell Beaker Czech Republic Knezeves [RISE566 / I4145] M 2279–2033 calBCE


123001 R1b1a1a2

[M269]

R1b1a1a2:L1353:19179540G->A; R1b1a1a2:PF6430:8070532T->A; R1b1a1a:CTS11985:23403749G->A; R1b1a:A702:10038192G->A; R1b1a:PF6263:21159055C->A; R:F63:7177189G->A H 263G, 750G, 1438G, 3107T, 5492C, 15326G, 16519C Allentoft 2015; Mathieson 2015; Olalde 2017
Bell Beaker Czech Republic Knezeves [RISE567] F
337289

U5b2c 73G, 150T, 263G, 723G, 750G, 1438G, 1721T, 1955A, 2706G, 3107T, 3197C, 7970A, 7977A, 9477A, 11467G, 11719A, 12308G, 12372A, 13017G, 13617C, 13637G, 14182C, 14766T, 15326G, 16192T, 16270T Allentoft 2015; Olalde 2017
Bell Beaker England Dairy Farm, Willington, (Bedfordshire) [I2452 / BEDFM2009.12, feature F.66 skeleton 186] M 2277–1920 calBCE 913255 R1b1a1a2a1a2c1

[L21 > DF13]

R1b1a1a2a1a2c1:CTS8221:17885577C->T; R1b1a1a2a1a:L52:14641193C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6497:21222868C->G; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6498:21312064C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1:CTS2134:14193384G->A; R1b1:M415:9170545C->A; R1b:M343:2887824C->A; etc

H1e1a Olalde 2017
Battle Axe/ Nordic LN Sweden Lilla Beddinge 56 [RISE98] M 2275-2032 BC 1,125,643

R1b1a1a2a1a1 M405 / S21/ U106 K1b1a1 73G, 152C, 263G, 750G, 1438G, 1811G, 2706G, 3107T, 3480G, 5913A, 7028T, 9055A, 9698C, 9962A, 10289G, 10398G, 10550G, 11299C, 11467G, 11719A, 11923G, 12308G, 12372A, 13967T, 14167T, 14766T, 14798C, 15257A, 15326G, 15946T, 16093C, 16224C, 16311C, 16319A, 16463G, 16519C Allentoft 2015; Mathieson 2015
Scotland Dryburn Bridge, East Lothian [I2567 / GENSCOT14, Burial 5, Cist 1] M 2275–1884 calBCE 589224 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a2:PF6505:21784286G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS8436:18026855G->A; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L780:21183643A->G; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; R1:CTS2565:14366723C->T; etc. U4a1a Olalde 2017
Bell Beaker Germany Unterer Talweg 58-62, Augsburg [E09538 / Feature 68 Skeleton 1] M 2471–2300 calBCE 545714 G2a2a1a2a1a G2a2a1a2a1a:FGC5671^:7652689G->A; G2a2a1a2a1a:Z6219:15894131C->T; G2a2a1a2a1:PF3239:17317628C->T; G2a2a1a2a1:PF3240:17589665C->T; G2a2a1a2a1:Z6130:6785680A->G; G2a2a1a2a:PF3238:17256431G->A; G2a2a1a2:L91:21645555G->C; G2a2a:PF3147:7738069G->A; G2a2a:PF3150:8476569T->C; G2a2a:PF3159:14815695C->G; G2a2a:PF3166:16735582T->G; G2a2a:PF3185:22894488C->T; G2a:CTS6753:17090976C->T; G2a:F2274:16802506G->T; G2a:L31:14028148C->A; G2a:M3408:22109159G->C; G2a:P15:23244026C->T; G2a:PF3141:23973594T->G; G2:CTS10089:19248446G->A; G2:F1189:8427005A->G; G2:F1239:8482393C->T; G2:F1294:8545324T->A; G2:F1393:8719593G->A; G2:F3198:21401188G->T; G2:F3220:21637589G->C; G2:F3344:22697266G->A; G2:F3536:23768744C->T; G2:M3488:8687693T->A; G2:M3579:21147058A->G; G:CTS282:2871867A->G; G:CTS692:6955839A->G; etc J1c Olalde 2017
Bell Beaker Germany Unterer Talweg 58-62, Augsburg [E09569 / Grave I/3] M 2397–2149 calBCE 810078 R1b1a1a2a1a2b1

[U152 > L2]

R1b1a1a2a1a2b1:L2:5755550C->T; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:L1353:19179540G->A; R1b1a1a2:M520:4446430T->A; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a2:PF6509:22190371A->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:P297:18656508G->C; R1b1a1a:PF6463:16183412C->A; R1b1a1a:PF6498:21312064C->A; R1b1a:A702:10038192G->A; R1b1a:CTS3063:14637352T->C; R1b1a:CTS4244:15510064T->G; R1b1a:FGC36:13822833G->T; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1a:L1345:21558298G->T; R1b1:CTS910:7081561C->T; R1b1:CTS3625:15080010C->T; R1b1:L278:18914441C->T; R1b1:L1349:22722580T->C; R1b1:M415:9170545C->A; R1b:M343:2887824C->A; R1:CTS2565:14366723C->T; etc J1c2 Olalde 2017

Spain Nerja, Málaga [NE-NAP]
2260 BC

H11a or L1b 16129A, 16264T, 16270T, 16278T, 16293G, 16311C Fernández 2005
Corded Ware Germany Karsdorf [KAR 53]
2260-2203 BC

K2a5 C195T A247G C324T A16129G T16187C C16189T T16223C T16224C G16230A T16278C Brandt 2013

Spain Abauntz, Navarra [ABb1.1] M? 2240 BC

H? CRS Fernández 2005

Spain Abauntz, Navarra [AB23C]
2240 BC

R or H? 16126C, 16311C Fernández 2005

Spain Abauntz, Navarra [AB25E]
2240 BC

H14? 16256T Fernández 2005
Bell Beaker Germany Karsdorf [KAR 5]



U5b A16129G T16187C C16189T C16192T T16223C G16230A C16270T T16278C C16311T Brandt 2013
Bell Beaker England Over Narrows, Needingworth Quarry, Cambridgeshire [I2454 / OVE08 feature F.1080 skeleton 5487] F 2200–1980 calBCE 818929 R1a1a Olalde 2017
Bell Beaker England Windmill Fields, Ingleby Barwick (Stockton-on-Tees, County Durham) [I1767 / Sk2] M 2200–1970 calBCE 729987 I2a2a1a1a I2a2a1a1a:L1195:18865320G->A; I2a2a1:CTS9183:18732197A->G; I2a2a:P221:8353707C->A; I2a2:L35:22725379C->A; I2a2:L37:17516123T->C; I2a2:L368:6931594C->T; I2a2:P217:7628484C->T; I2a2:P218:17493630T->G; I2:L68:18700150C->T; I2:PF3664:8567995G->A; I:CTS88:2723755G->A; I:CTS646:6926038T->A; etc

U5a1a1 Olalde 2017
Bell Beaker Italy Via Guidorossi (Parma) [I2478 / Tomb1] M 2200–1930 calBCE 622069 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6425:7762947T->C; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6509:22190371A->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS7941:17755905G->A; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:PF6459:15286480G->C; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:CTS8436:18026855G->A; R1b1a:FGC35:18407611C->T; R1b1a:FGC36:13822833G->T; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1a:PF6249:8214827C->T; R1b1a:PF6263:21159055C->A; R1b1:CTS2134:14193384G->A; R1b1:L278:18914441C->T; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS2565:14366723C->T; etc K1a2a Olalde 2017
Bell Beaker England Amesbury (Boscombe) Down, Wiltshire [I2566 / 13385] M
2210–2030 calBCE 485091 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:L754:22889018G->A; R1b1:CTS2134:14193384G->A; R1b1:L1349:22722580T->C; R1b1:M415:9170545C->A; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; etc K1b1a1 Olalde 2017
Bell Beaker England Trumpington Meadows, Cambridge [I3256 / TRM10, skeleton [3384] second-degree relative of I3255 M 2204–2029 calBCE 843389 R1b1a1a2a1a2c1

[L21 > DF13]

R1b1a1a2a1a2c1:CTS8221:17885577C->T; R1b1a1a2a1a2c:L21:15654428C->G; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:M269:22739367T->C; R1b1a1a2:M520:4446430T->A; R1b1a1a2:PF6399:2668456C->T; R1b1a1a2:PF6409:4352151G->A; R1b1a1a2:PF6425:7762947T->C; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a2:PF6505:21784286G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6451:14116584T->A; R1b1a1a:PF6463:16183412C->A; R1b1a1a:PF6524:23452965T->C; R1b1a1:L388:17400785G->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:CTS8436:18026855G->A; R1b1a:FGC35:18407611C->T; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1a:L1345:21558298G->T; R1b1a:PF6263:21159055C->A; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L822:7960019G->A; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; etc

T2b Olalde 2017
Bell Beaker Germany Benzingerode-Heimburg [BZH 7 and 12]
2201-2136 BC

U5a1a2a 2 samples of A16129G T16187C C16189T T16223C G16230A C16256T C16270T T16278C C16311T A16399G Brandt 2013; Haak 2015
Bell Beaker Denmark Damsbo [D 1]
2200 BC

U4 7028T, 12308G, 16356C Melchior 2010
Bell Beaker Denmark Damsbo [D 2]
2200 BC

U5a2a? 7028T, 12308G, 16114A, 16192T, 16256T, 16270T, 16294T, reported as U5a Melchior 2010
Bell Beaker Germany Benzingerode-Heimburg [BZH 5]
2198-2161 BC

T2a1b T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T C16311T T16324C Brandt 2013
Bell Beaker Germany Eulau [EUL 22]
2198-2162 BC

W A16129G T16187C C16189T G16230A T16278C C16292T C16311T Brandt 2013
Bell Beaker Germany Eulau [EUL 31]



T1a T16126C A16129G T16140C A16163G C16186T T16187C T16223C G16230A T16278C C16294T C16311T Brandt 2013

Denmark Falshoj [RISE71] M 2196-2023 BC 203,604



H3b 263G, 750G, 1438G, 2581G, 3107T, 15326G, 16519C Allentoft 2015
Bell Beaker England Amesbury (Boscombe) Down, Wiltshire [I2598 / 12134] M
2140–1940 calBCE 85900 R1b1a1a2a1a

[P311]

R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:PF6497:21222868C->G; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1:CTS2565:14366723C->T; R:P227:21409706G->C H Olalde 2017
Bell Beaker England Trumpington Meadows, Cambridge [I3255 / TRM10, skeleton 3383] second-degree relative of I3256 F 2136–1951 calBCE 861509 T2b

Olalde 2017
Bell Beaker England Over Narrows, Needingworth Quarry, Cambridgeshire [I2455 / OVE08, feature F.1079 skeleton 5486] F 2130–1910 calBCE 120698 U5a2c3a Olalde 2017
Baltic LN Lithuania Spiginas [2] M 2130-1750 cal BCE 870598 R1a1a1b I4a Mittnik 2017

Spain Tres Montes, Navarra [TM-3]
2130 BC



Fernández 2005

Spain Tres Montes, Navarra [TM-6]
2130 BC

H or K 16224C Fernández 2005

Spain Tres Montes, Navarra [TM-8]
2130 BC

R or H? 16126C, 16311C Fernández 2005

Spain Tres Montes, Navarra [TM-11]
2130 BC


16261T, 16278T, 16294T, 16309G Fernández 2005

Spain Tres Montes, Navarra [TM-18]
2130 BC

L2 16261T, 16278T 16294T, 16309G Fernández 2005
Bell Beaker Germany Karsdorf [KAR 52 ]
2083-2042 BC

U4c1 C146T C152T A247G A16129G C16179T T16187C C16189T T16223C G16230A T16278C C16311T T16356C Brandt 2013
Bell Beaker? Germany Quedlinburg [QUEXII 2]
2050-1940 BC

H4a1 G73A C146T C152T C195T A247G A769G A825t A1018G G2706A A2758G C2885T T3594C C3992T A4024G G4104A T4312C T5004C T7028C G7146A T7256C A7521G T8468C T8655C G8701A G9123A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C C14365T A14582G T14766C A16129G T16187C C16189T T16223C G16230A T16278C C16311T C16519T Adler 2012; Brotherton 2013; Brandt 2013
Bell Beaker England Amesbury (Boscombe) Down, Wiltshire [I2460 / 62027] F
2030–1820 calBCE 23341 T2a1a Olalde 2017
Nordic LN Sweden Abekas I [RISE179] M 2010-1776 BC 50,762

I CTS674, CTS1301, FI3, CTS10941 K1a3 73G, 497T, 750G, 1438G, 2019A, 2706G, 9055A, 10398G, 11299C, 11467G, 13117G, 14798C, 15326G, 16224C, 16519C Allentoft 2015; Mathieson 2015
Corded Ware Estonia Ardu, Harju [2] M 4,871–4,505 cal BP R1a1a1b Z645 U5b2c Saag 2017
Corded Ware Estonia Ardu, Harju [1] M 4,800–4,000 cal BP R1a1a1b Z645 T2a1a Saag 2017
Poltavka Russia Lopatino II, Sok River, Samara [I0440 / SVP 53] M 2885-2665 BC 1,058,753

R1b1a1a2a2

[CTS1078 / Z2103]

CTS1078, Z2105 I3a Mathieson 2015
Poltavka Russia Grachevka II, Sok River, Samara [I0371 / SVP 11] M 2875-2580 BC 613,900 R1b1a1a2 [M269] L773, L265, F1794, PF6475, L500, PF6482, PF6495, PF6505, PF6509, CTS12478 U2d2 Mathieson 2015
Poltavka Russia Kutuluk III, Kutuluk River, Samara [I0126 / SVP 51] M 2867-2484 BC 339,812 R1b1a1a2a2

[CTS1078 / Z2103]

Z2105 H6a2 Mathieson 2015
Poltavka Russia Nikolaevka III, Samara River, Samara [I0374 / SVP 16] M 2800-2200 BC 891,602 R1b1a1a2a L23 H13a1a Mathieson 2015; Lazaridis 2016
Potapovka Russia Utyevka VI, Samara River, Samara [I0246 / SVP 41] M 2469-1928 BC 90,021 P1 P284 C1 Mathieson 2015; Lazaridis 2016
Minoan Greece Ayios Charalambos cave, Lassithi plateau, Crete [6AH]
4400-3700 BP

H 16266T Hughey 2013
Minoan Greece Ayios Charalambos [7AH]
4400-3700 BP

H 2706A, (no 16519) Hughey 2013
Minoan Greece Ayios Charalambos [16AH, 17AH, 18AH, 20H]
4400-3700 BP

H 4 samples 2706A Hughey 2013
Minoan Greece Ayios Charalambos [7H]
4400-3700 BP

H 2706A, 16290T Hughey 2013
Minoan Greece Ayios Charalambos [8H]
4400-3700 BP

H 2706A, 16172C Hughey 2013
Minoan Greece Ayios Charalambos [#11]
4400-3700 BP

H 16290T, 16304C, 16519T Hughey 2013
Minoan Greece Ayios Charalambos [9AH]
4400-3700 BP

H5 2706A, 16304C Hughey 2013
Minoan Greece Ayios Charalambos [12 AH]
4400-3700 BP

H7 2706G Hughey 2013
Minoan Greece Ayios Charalambos [2AH]
4400-3700 BP

H13a1a 14766C Hughey 2013
Minoan Greece Ayios Charalambos [1AH]
4400-3700 BP

HV 14766C, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [24H]
4400-3700 BP

HV 11719A, 14766C, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [#6]
4400-3700 BP

HV 2706G, 7028T, 11719A, 14766C, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [#12]
4400-3700 BP

I5 16129A, 16148T, 16186T, 16223T Hughey 2013
Minoan Greece Ayios Charalambos [#13]
4400-3700 BP

I5 199T, 250C, 16129A, 16148T, 16223T, 16391A Hughey 2013
Minoan Greece Ayios Charalambos [#17]
4400-3700 BP

I5 10034C, 16129A, 16148T, 16186T, 16223T, 16391A Hughey 2013
Minoan Greece Ayios Charalambos [26H]
4400-3700 BP

J2 13708A, 16069T, 16126C, 16232T, 16291T, 16293C Hughey 2013
Minoan Greece Ayios Charalambos [11AH]
4400-3700 BP

K 12372A, 16111T, 16185T, 16224C, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [15AH]
4400-3700 BP

K 3480G, 14167T, 16111T, 16185T, 16224C, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [11AH]
4400-3700 BP

K 12372A, 16111T, 16185T, 16224C, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [#14]
4400-3700 BP

K 12372A, 16111T, 16224C, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [#21]
4400-3700 BP

K 11467G, 12308G, 16224C, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [#22]
4400-3700 BP

K 10550G, 11299C, 11467G, 12308G, 16224C, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [#7]
4400-3700 BP

R0 73A, 2706G, 7028T Hughey 2013
Minoan Greece Ayios Charalambos [18AH]
4400-3700 BP

T 1888A, 16126C, 16294T, 16296T Hughey 2013
Minoan Greece Ayios Charalambos [9H]
4400-3700 BP

T1 10463C, 15928A, 16126C, 16163G, 16186T, 16189C, 16263C,16294T, 16296T Hughey 2013
Minoan Greece Ayios Charalambos [3AH]
4400-3700 BP

T2 4917G, 10463C, (no 15607), 16126C, 16294T, 16296T, 16304C Hughey 2013
Minoan Greece Ayios Charalambos [10AH]
4400-3700 BP

T2 14905A, 16126C, 16294T, 16296T Hughey 2013
Minoan Greece Ayios Charalambos [#19]
4400-3700 BP

T2 13368A, 16089A, 16126C, 16294T, 16296T, 16304C Hughey 2013
Minoan Greece Ayios Charalambos [#23]
4400-3700 BP

T3 4917G, 11251G, 15607G, 15928A, 16126C, 16294T, 16296T, 16339T Hughey 2013
Minoan Greece Ayios Charalambos [21H]
4400-3700 BP

T5 10463C, 13368A, 15607G, 15298A, 16126C, 16153A, 16240A?C, 16294T, 16296T Hughey 2013
Minoan Greece Ayios Charalambos [#4]
4400-3700 BP

U 16182A?C, 16183A?C, 16189C, 16234T, 16290T, 16324C Hughey 2013
Minoan Greece Ayios Charalambos [4H]
4400-3700 BP

U5a 12308G, 12372A, 16192T, 16256T, 16270T, 16311C Hughey 2013
Minoan Greece Ayios Charalambos [6H]
4400-3700 BP

W 204C, 207A, 15884C, 16093C, 16223T, 16243C, 16292T Hughey 2013
Minoan Greece Ayios Charalambos [#8]
4400-3700 BP

X 153G, 14470C, 16145A, 16179T, 16189C, 16223T, 16278T, 16332T, 16344T, 16362C Hughey 2013
Okunevo Russia Verkhni Askiz Village [RISE515] F 2340-2145 BC

A8a 64T, 73G, 146C, 235G, 263G, 663G, 722T, 1438G, 1736G, 4248C, 4824G, 11719A, 12705T, 14766T, 15326G, 16242T, 16290T, 16293C, 16319A Allentoft 2015
Bell Beaker Netherlands Oostwoud-Tuithoorn, West Frisia [I4068 / skeleton 228-M3] M 2300–1900 BCE 469691 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:PF6459:15286480G->C; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:CTS8436:18026855G->A; R1b1a:L754:22889018G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b:M343:2887824C->A; R1:CTS3321:14829196C->T; R1:CTS5611:16394489T->G; etc

U5a2a1 Olalde 2017
Bell Beaker Netherlands Oostwoud-Tuithoorn, West Frisia [I4076 / skeleton 247-M18] F 2300–1900 BCE 436123 K1d Olalde 2017
Sintashta Russia Bulanovo [RISE386] M 2298-2045 BC 315,156

R1a

[*R1a1a1b2a2]

L62. [*Tagankin reads Z2124+, and downstream SNPs Z2125+, S23592+, YP1456+] J1c1b1a 73G, 185A, 228A, 263G, 295T, 462T, 482C, 489C, 1438G, 2706G, 3010A, 3107T, 3394C, 5773A, 7184G, 10398G, 10463C, 11251G, 11719A, 12612G, 13708A, 14766T, 14798C, 15326G, 15452A, 16069T, 16126C Allentoft 2015; Mathieson 2015; *additional info from Vladimir Tagankin
Corded Ware/ Proto-Unetice Poland Leki Male [RISE431] M 2286-2048 BC 124,341 K(xLT)

M526 T2e 73G, 150T, 263G, 750G, 2706G, 3107T, 6359G, 8155A, 8697A, 11251G, 11719A, 11812G, 14623T, 14766T, 14905A, 15326G, 15452A, 15607G, 15613G, 15928A, 16126C, 16153A, 16294T, 16296T, 16519C Allentoft 2015; Mathieson 2015

Spain El Portalón, Sierra de Atapuerca [ATP20] F 4239-4000 cal BP

U5a1c 146T, 152T, 153G, 247G, 769G, 825T, 1018G, 2758G, 2885T,
3197C, 3594C, 4312C, 7256C, 8468C, 8655C, 8701A, 9477A,
9540T, 10398A, 10664C, 10688G, 10810T, 10873T, 10915T, 11467G, 11914G, 12308G, 12372A, 12705C, 13105A, 13276A, 13506C, 13617C, 13650C, 13802T, 16129G, 16187C, 16189T,
16192T, 16223C, 16230A, 16256T, 16278C, 16320T, 16399G, 195T, 7013A, 14766C, 16519T
Günther 2015
Bell Beaker Netherlands Oostwoud-Tuithoorn, West Frisia [I4074 / skeleton 242-M14] M 2278–1914 calBCE 775215 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L11:17844018T->C; R1b1a1a2a1a:L52:14641193C->T; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS8665:18137831T->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6451:14116584T->A; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:CTS8436:18026855G->A; R1b1a:CTS8612:18109555C->A; R1b1a:FGC36:13822833G->T; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1a:PF6263:21159055C->A; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L780:21183643A->G; R1b1:L1349:22722580T->C; R1b1:M415:9170545C->A; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; etc H Olalde 2017
Unetice Germany Esperstedt [ESP 29 / I0117] F 2272-2039 calBCE (3743±25 BP, MAMS-21496) 874,673

I3a T16086C T16187C C16189T G16230A T16278C C16311T G16391A Adler 2012; Brandt 2013; Haak 2015; Lazaridis 2016
Okunevo Russia Verkhni Askiz Village [RISE516] F 2201-2036 BC

H6a1b 239C, 263G, 750G, 1438G, 3107T, 3915A, 9380A, 10589A, 15326G, 16362C, 16482G Allentoft 2015
Bronze Age Germany Untermeitingen [RISE471] M
116,341

R1b1a2a1a
P310
J1c1b 73G, 185A, 228A, 263G, 295T, 462T, 482C, 489C, 750G, 1438G, 2706G, 3010A, 3107T, 3394C, 7184G, 10398G, 11251G, 11719A, 12612G, 12810G, 13708A, 14766T, 14798C, 15326G, 15452A, 16069T, 16126C Allentoft 2015; Mathieson 2015
Unetice Germany Eulau [EUL 51]
2200-2136 BC

U2e1f A16051G A16129c T16187C T16223C G16230A T16278C T16362C Brandt 2013
Potapovka Russia Utyevka VI, Samara River, Samara [I0419 / SVP 27] M 2200-1900 BC 474,599 R1a1a1b S441 U2e1h Mathieson 2015
Unetice Germany Benzingerode-Heimburg [BZH 14]
2200-1575 BC

H82a G73A C146T C152T A247G A769G A825t A1018G G2706A A2758G C2885T T3594C G4104A T4312C T7028C G7146A T7256C A7521G T8468C T8655C G8701A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C T14766C A16129G T16187C C16189T A16220G T16223C G16230A T16278C C16311T Brotherton 2013; Brandt 2013
Unetice? Germany Benzingerode-Heimburg [BZH 1]
2200-1550 BC

H11a G73A C146T C152T A247G A769G A825t T961g A1018G G2706A A2758G C2885T T3594C G4104A T4312C T7028C G7146A T7256C A7521G T8448C T8468C T8655C G8701A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C G13759A T14766C A16129G T16187C C16189T T16223C G16230A T16278C A16293G C16519T Brotherton 2013; Brandt 2013
Unetice Germany Eulau [EUL 35]
2200-1550 BC

H A16129G T16187C T16223C G16230A T16278C T16304C C16311T Brandt 2013
Unetice Germany Eulau [EUL 41]
2200-1550 BC

H4a1a1a C146T C152T C195T A247G A769G A825t A1018G G2706A A2758G C2885T T3594C C3992T A4024G G4104A T4312C T5004C T7028C G7146A T7256C A7521G G8269A T8468C T8655C G8701A G9123A C9540T A10044G G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C C13545T T13650C C14365T A14582G T14766C A16129G T16187C C16189T T16223C G16230A T16278C C16311T C16519T Brotherton 2013 ; Brandt 2013; Mathieson 2015
Unetice Germany Eulau [EUL 42]
2200-1550 BC

K C195T A247G A16129G T16187C C16189T T16223C T16224C G16230A T16278C Brandt 2013
Unetice? Germany Eulau [EUL 48]
2200-1550 BC

K1a T16093C A16129G T16187C T16223C T16224C G16230A T16278C Brandt 2013
Unetice? Germany Eulau [EUL 50]
2200-1550 BC

W T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T T16304C C16311T Brandt 2013
Unetice Germany Eulau [EUL 59]
2200-1550 BC

HV6 A16129G T16172C T16187C C16189T T16223C G16230A T16278C Brandt 2013
Unetice? Germany Karsdorf [KAR 23]
2200-1550 BC

H A16129G T16187C C16189T T16223C G16230A T16278C C16291T C16311T Brandt 2013
Unetice? Germany Karsdorf [KAR 35]
2200-1550 BC

T2 315.1C C146T C152T C195T A247G T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T C16296T C16311T Brandt 2013
Unetice Germany Karsdorf [KAR 44]
2200-1550 BC

H G73A C146T C152T C195T A247G T252C 315.1C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Unetice Germany Karsdorf [KAR 48]
2200-1550 BC

U5a1 C146T C152T C195T A247G 315.1C A16129G T16187C C16189T C16192T T16223C G16230A C16256T C16270T T16278C C16311T A16399G Brandt 2013
Unetice Germany Leau 2 [LEA 2 and 3]
2200-1550 BC

V 2 samples G73c C146T C152T C195T A247G 309.1C 309.2C 315.1C A16129G T16187C C16189T T16223C G16230A T16278C T16298C C16311T Brandt 2013
Unetice Germany Leau 2 [LEA 4]
2200-1550 BC

H6a1b3 G73A C146T C152T C195T T204C T239C A247G 309.1C 315.1CA16129G T16187C C16189T C16193T A16219G T16223C G16230A T16278C C16311T T16362C Brandt 2013
Unetice Germany Plotzkau 3 [PLOTZ 1]
2200-1550 BC

J2b1a C146T C150T C195T A247G C295T 315.1C C16069T T16126C A16129G T16187C C16189T C16193T T16223C G16230A C16311T Brandt 2013
Unetice Germany Plotzkau 3 [PLOTZ 2 and 3]
2200-1550 BC

H 2 samples of G73A C146T C152T C195T A247G 315.1C A16129G T16187C C16189T T16223C G16230A T16278C Brandt 2013
Unetice Germany Plotzkau 3 [PLOTZ 4, 5 and 8]
2200-1550 BC

U5a 3 samples of C146T C152T C195T A247G 315.1C A16129G T16187C C16189T C16192T T16223C G16230A C16256T C16270T T16278C C16311T Brandt 2013
Unetice Germany Plotzkau 3 [PLOTZ 6 and 7]
2200-1550 BC

I1a1 2 samples of C146T C152T C195T T199C G203A T204C A247G T250C 315.1C T16172C T16187C C16189T G16230A T16278C G16391A Brandt 2013
Unetice? Germany Quedlinburg VII 2 [QLB 29]
2200-1550 BC

U8a1a A16129G A16146G T16187C C16189T T16223C G16230A T16278C C16311T T16342C Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 32]
2200-1550 BC

R A16129G T16187C C16189T T16223C G16230A T16278C T16362C Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 33]
2200-1550 BC

X A16129G T16187C G16230A C16311T Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 34]
2200-1550 BC

U2e1f A16051G T16086C A16129c T16187C T16223C G16230A T16278C T16362C Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 35 and 43]
2200-1550 BC

H5 2 samples of A16129G T16187C C16189T T16223C G16230A T16278C T16304C C16311T Brandt 2013
Unetice? Germany Quedlinburg VII 2 [QLB 36]
2200-1550 BC

J1c3f T16063C C16069T T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 37]
2200-1550 BC

U2e2 A16183c A16051G T16092C A16129c T16187C T16223C G16230A T16278C C16311T T16362C Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 40]
2200-1550 BC

J C16069T T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 42]
2200-1550 BC

J C16069T T16126C A16129G T16187C T16223C G16230A T16278C C16311T Brandt 2013
Unetice Germany Rocken [ROC 2 and 3]
2200-1550 BC

U8b1a1 2 samples of C152T C195T A247G 309.1C 309.2C 315.1C A16182c A16183c A16066G T16187C T16223C G16230A C16234T T16278C C16311T Brandt 2013
Unetice Germany Rocken [ROC 4]
2200-1550 BC

U2e1 309.1C 309.2C 315.1C C146T C195T T217C A247G C340T A16183c A16051G A16129c T16187C T16223C G16230A T16278C C16311T T16362C Brandt 2013
Unetice Germany Rocken [ROC 5]
2200-1550 BC

U4 A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Brandt 2013
Unetice Germany Rocken [ROC 7]
2200-1550 BC

H A16129G T16187C C16189T T16223C G16230A C16261T T16278C A16293t C16311T Brandt 2013
Unetice Germany Rocken [ROC 8]
2200-1550 BC

J1c2e C16069T T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16311T C16366T Brandt 2013
Unetice Germany Rocken [ROC 9]
2200-1550 BC

I T16187C C16189T G16230A T16278C C16311T G16391A Brandt 2013
Unetice Germany Rocken [ROC 10]
2200-1550 BC

U2e2 A16183c A16051G T16092C A16129c T16187C T16223C G16230A T16278C C16311T A16335G T16362C Brandt 2013
Unetice Germany Rocken [ROC 11]
2200-1550 BC

K1a T16093C A16129G T16187C C16189T T16223C T16224C G16230A T16278C Brandt 2013
Bell Beaker Netherlands Oostwoud-Tuithoorn, West Frisia [I4073 / skeleton 236-M13] M 2196–1903 calBCE 761432 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L52:14641193C->T; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS8728:18167403C->T; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:F1794:14522828G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a2:PF6509:22190371A->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6463:16183412C->A; R1b1a1:L388:17400785G->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:FGC35:18407611C->T; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1a:PF6263:21159055C->A; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L780:21183643A->G; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; R1:CTS2565:14366723C->T; etc U5a2b3 Olalde 2017
Nordic Late Neolithic Denmark Marbjerg [RISE42] M 2191-1972 BC

H3v+16093 263G, 408A, 750G, 1438G, 3107T, 15326G, 16093C, 16519C Allentoft 2015
Mako Hungary Kompolt-Kigyoser [I1502/BR 1] F 2190-1980 BC

K1c1
Gamba 2014; Mathieson 2015
Unetice Poland Polwica [RISE145] F 2188-1958 BC

H6a1b 239C, 750G, 1438G, 3107T, 3915A, 4288T, 9380A, 10090T, 10589A, 15326G, 16362C, 16482G Allentoft 2015
Bell Beaker Netherlands Oostwoud-Tuithoorn, West Frisia [I4069 / skeleton 229-M4] M 2188–1887 calBCE 721500 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L52:14641193C->T; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L500:18180446C->A; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6459:15286480G->C; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:FGC35:18407611C->T; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:L1345:21558298G->T; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; etc

U5a1a1 Olalde 2017
Yamna Ukraine Liubasha, kurgan 2 [L15] 2161 ± 107 BC C4a2 16218T, 16223T, 16298C, 16327T,16357C Nikitin 2017; Newton 2011
Unetice? Germany Esperstedt [ESP 27] F 2155-2034 BC

T2c T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16292T C16294T C16311T Adler 2012; Brandt 2013
Unetice Poland Chociwel [RISE139] M 2153-1923 BC

U2e1f1 489C, 508G, 3720G, 8155A, 11467G, 13734C, 15326G Allentoft 2015
Unetice Germany Quedlinburg VII 2 [QLB 30]
2150-1960 BC

K A16129G T16187C C16189T T16223C T16224C G16230A T16278C Brandt 2013
England Porton Down, Wiltshire [I2600 / 5108] daughter of I2457/13382 F
2140-1940 calBCE 524726
U5b1+16189 Olalde 2017
Scotland Eweford Cottages, East Lothian [I2569 / GENSCOT17] M 2140–1916 calBCE 23665 R1b1a1a2a1a2

[P312]


R1b1a1a2a1a2:P312:22157311C->A K1a3a Olalde 2017
Unetice Germany Eulau [I0804 / EUL 57] M
2137-1965 calBCE (3671±26 BP)

62,496 I2 L68+, P158+ H3 G73A C146T C195T A247GA769G A825t A1018G G2706A A2758G C2885T T3594C G4104A T4312C T6776C T7028C G7146A T7256C A7521G T8468C T8655C G8701A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C T14766C A16129G T16187C C16189T T16223C G16230A T16278C C16311T

T152C!

Brandt 2013; Haak 2015; Mathieson 2015; Lazaridis
Bell Beaker England Yarnton, Oxfordshire [I2445 / SK 8633 (YCF 95)] M 2137–1930 calBCE 854563 R1b1a1a2a1a2c1

[L21 > DF13]

R1b1a1a2a1a2c1:CTS8221:17885577C->T; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS8665:18137831T->C; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1a:CTS8436:18026855G->A; R1b1a:FGC36:13822833G->T; R1b1a:FGC41:7900883C->A; R1b1a:L754:22889018G->A; R1b1a:PF6249:8214827C->T; R1b1a:PF6263:21159055C->A; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L780:21183643A->G; R1b1:L822:7960019G->A; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; etc

X2b6 Olalde 2017
Maros Hungary Szoreg - C [RISE371] F 2136-1941 BC 53,698

U5a2b 73G, 750G, 1438G, 3107G, 3197C, 9477A, 9548A, 11467G, 12308G, 12372A, 13617C, 14344G, 14766T, 14793G, 15326G, 16192T, 16256T, 16270T, 16526A Allentoft 2015
England Reaverhill, Barrasford, Northumberland [I2618 / MOA 1964.2 Box 102A] M 2135–1951 calBCE 643010 R1b1a1a2a1a

[L151]

R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS3575:15037433C->G; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L500:18180446C->A; R1b1a1a2:L773:7220727A->G; R1b1a1a2:M520:4446430T->A; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS7941:17755905G->A; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS3063:14637352T->C; R1b1a:FGC35:18407611C->T; R1b1a:L754:22889018G->A; R1b1a:L1345:21558298G->T; R1b1a:PF6249:8214827C->T; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1:CTS997:7132713G->A; R1:CTS2565:14366723C->T; etc. H1bs Olalde 2017
Unetice Czech Republic Velke Prilepy [RISE577] F
738,363



T2b 73G, 263G, 709A, 930A, 1438G, 2706G, 3107T, 4917G, 8697A, 10463C, 11251G, 11719A, 11812G, 14233G, 14766T, 14905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16294T, 16296T, 16304C, 16519C Allentoft 2015
Unetice Czech Republic Moravska Nova Vez [RISE586] F
195,448



K1b1a 73G, 152C, 263G, 750G, 1438G, 9055A, 9962A, 10289G, 10398G, 10550G, 11299C, 11467G, 11719A, 11923G, 12308G, 12372A, 13759A, 14167T, 14766T, 14798C, 15257A, 15326G, 15946T, 16093C, 16224C, 16311C, 16319A, 16463G, 16519C Allentoft 2015
Remedello Italy Remedello di Sotto [RISE486] M 2134-1773 BC 203,575

I [*I2a1a1] PF3640, PF3661, FGC2415, CTS2536, CTS4272, PF3800, CTS10941, CTS11540; [*Tagankin reads Y3995+ PF4051+ PF4047+ PF3986+ PF3848+ CTS7671/PF4026+ CTS5526/PF3741+ PF4055+ PF4049+ CTS9771/PF4040+ V3296/CTS8968/PF4036+ CTS8830/PF4035+ CTS7977/PF4028+ CTS2925/PF3710+ L1300/PF4050+ Z2053/CTS11766/PF4071+, Y11895+ Y11222+ Y11223+ Z27401/CTS7023/PF6937+, Z27397+ Y15658- , Y15463- Y15461- I-L160: PF4066- CTS11972/PF4072- CTS6406/PF4093- ] J1c1b 73G, 185A, 228A, 263G, 295T, 462T, 482C, 489C, 750G, 1438G, 2594C, 2706G, 3107C, 3394C, 7184G, 8006T, 9490T, 10398G, 11251G, 11719A, 12612G, 13436T, 13708A, 14429T, 14766T, 14798C, 15326G, 15452A, 16069T, 16126C Allentoft 2015; Mathieson 2015; *additional info from Vladimir Tagankin
Unetice? Germany Esperstedt [I0116/

ESP 4]

M 2134-1939 calBC (3650±32 BP) 640,413 I2c PF3827+, L597+, M438+ W3a1 A16129G C16147g T16187C C16189T G16230A T16278C C16292T C16311T Adler 2012; Brandt 2013; Haak 2015; Mathieson 2015; Lazaridis 2016
Unetice Germany Eulau [EUL 47]
2133-2080 BC

K2 C152T C195T A247G A16129G T16187C C16189T T16223C T16224C G16230A T16278C Brandt 2013
Unetice Germany Eulau [EUL 41 / I0803] F 2132-1942 calBCE (3650±26 BP)

224,184

H4a1a1 C146T C152T C195T A247G 309.1C 315.1C A769G A825t A1018G G2706A A2758G C2885T T3594C C3992T A4024G G4104A T4312C T5004C T7028C G7146A T7256C A7521G G8269A T8468C T8655C G8701A G9123A C9540T A10044G G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C C13545T T13650C C14365T A14582G T14766C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013; Haak 2015; Lazaridis 2016
Unetice Germany Esperstedt [I0114/ESP 2] M 2131-1979 BC I2a2b L368+, L181+, P218+, P217+, M438+, L34-, P223-, M223-, [L39+] I3a T16086C T16187C C16189T G16230A T16278C C16311T G16391A Adler 2012; Brandt 2013; Haak 2015; Mathieson 2015; [Additional info on SNP L39 from Hans De Beule and Atanas Kumbarov]
Vatya Hungary Szazhalombatta-Foldvar [RISE254] M 2128-1909 BC 42,943

I [*I2a1] FI2, CTS2514, CTS3384

[*Tagankin Z2657/CTS7669/PF3765+ Z2673/CTS12003/PF3846+ Z2652/CTS4568/PF3733+, FGC18615/Y12523- FGC18580/Y7862- FGC18589/Y5341- S6669-, Z2616/S5140+ V1191/Z2611/PF3638+]

J1c9 73G, 185A, 263G, 462T, 489C, 750G, 1438G, 1462A, 1738C, 2706G, 3010A, 3107T, 6887T, 7028T, 10398G, 11251G, 11323T, 11719A, 12612G, 13708A, 14766T, 14798C, 15326G, 15452A, 16069T, 16126C Allentoft 2015; Mathieson 2015; *additional info from Vladimir Tagankin
Unetice Germany Karsdorf [KAR 49]
2126-2084 BC

H5 A16129G T16187C C16189T T16223C G16230A T16278C T16304C C16311T Brandt 2013
Sintashta Russia Stepnoe VII [RISE392] M 2126-1896 BC 526,675

R1a1a1b

[*R1a1a1b2a2a]

S224 [*Tagankin reads Z2123 +, and downstream SNPs Y877+, Y939-] J2b1a2a 73G, 150T, 152C, 235G, 263G, 295T, 489C, 750G, 1438G, 2706G, 3107T, 5633T, 6216C, 6893T, 7028T, 7476T, 8962G, 10172A, 10398G, 11251G, 11719A, 12612G, 13708A, 14766T, 15212G, 15257A, 15326G, 15452A, 15812A, 16069T, 16126C, 16193T, 16278T, 16519C Allentoft 2015; Mathieson 2015; *additional info Vladimir Tagankin
Potapovka Russia Utyevka IV, Samara River, Samara [I0418 / SVP 24] F 2125-2044 BC 138,537

T1a1

Mathieson 2015
Sintashta Kazakhstan Tanabergen II [RISE391] F 2120-1887 BC 55,088



N1a1a1a1 73G, 152C, 199C, 204C, 263G, 669C, 1438G, 1719A, 2706G, 3107T, 3336C, 5315G, 7028T, 8164T, 8901G, 9300A, 10238C, 10398G, 11719A, 12501A, 12705T, 13780G, 14766T, 15043A, 15326G, 16147A, 16172C, 16189C, 16223T, 16248T, 16320T, 16355T, 16519C Allentoft 2015
Unetice Germany Halberstadt [I0047 / HAL 16] F 2111-1891 calBCE (3612±30 BP)

836,247

V9 A16129G T16187C C16189T T16223C G16230A T16278C T16298C C16311T Brandt 2013; Haak 2015; Mathieson 2015; Lazaridis 2016
Thracian Romania South East [1BA]
2100-1200 BC

H17 16129C Cardos 2004
Thracian Romania South East [5BA]
2100-1200 BC

? 16186T, 16189C 16299T Cardos 2004
Thracian Romania South East [7BA]
2100-1200 BC

? 16129A, 16223T Cardos 2004
Thracian Romania South East [8BA]
2100-1200 BC

H22? 16145A Cardos 2004
Thracian Romania South East [9BA]
2100-1200 BC

HV? 16311C Cardos 2004
Unetice Germany Karsdorf [KAR 42]
2083-2042 BC

I C146T C152T C195T T199C T204C A247G T250C 309.1C 315.1C T16187C C16189T G16230A T16278C C16311T G16391A Brandt 2013
Unetice Germany Quedlinburg [QUEVIII 4]
2050-1800 BC

H7h A4793G, G16213A Adler 2012; Brotherton 2013
Unetice Germany Esperstedt [ESP 6]
2050-1800 BC

I1 T16187C C16189T G16230A G16391A Adler 2012; Brandt 2013
Unetice Germany Esperstedt [ESP 23]
2050-1800 BC

U5a2a C16114a A16129G T16187C C16189T C16192T T16223C G16230A C16256T C16270T T16278C C16294T C16311T Brandt 2013
Unetice Germany Quedlinburg [QUEVIII 3] M 2050-1800 BC

T1 T16126C, A16163G, C16186T, T16189C, C16294T Adler 2012
Unetice Germany Quedlinburg [QUEXIV 1] F 2050-1800 BC

T2 T16126C, 16292T, 16294T Adler 2012
Unetice Germany Quedlinburg [QUEVIII 2] F 2050-1800 BC

U2 A16126C, A16163G, G16129c, A16183c, T16189C, T16362C Adler 2012
Unetice Germany Quedlinburg [QUEVIII 5]
2050-1800 BC

U5a1a 16256T, 16270T, a16399G Adler 2012
Unetice Germany Quedlinburg [QUEXII 7] F 2050-1800 BC

U5a1a C16192T, C16222T, C16256T, C16270T, a16399G Adler 2012
Unetice Germany Esperstedt [ESP 12 and 31] F 2050-1800 BC

X Two samples A16129G T16187C G16230A C16311T Adler 2012; Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 41]
2040-1910 BC

W3a1 C146T C152T A189G C194T T204C G207A A247G C146T C152T A189G C194T T204C G207A A247G 309.1C 315.1C A16129G T16187C C16189T G16230A T16278C C16292T C16311T Brandt 2013
Middle Bronze Age Hungary Battonya Voris Oktober [RISE349] F 2034-1748 BC 96,499

T2b3 73G, 183G, 195C, 263G, 709A, 750G, 930A, 1438G, 2706G, 3107T, 4917G, 8697A, 10463C, 10750G, 11251G, 11719A, 11812G, 14233G, 14766T, 14905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16169T, 16189C, 16294T, 16296T, 16304C, 16519C Allentoft 2015
Unetice? Germany Eulau [EUL 28]
2034-1954 BC

J1b1a1 C16069T T16126C A16129G G16145A T16172C T16187C C16189T C16222T T16223C G16230A C16261T T16278C C16311T Brandt 2013
Unetice Germany Eulau [EUL 57A]
2030-1987 BC

U5 A16129G T16187C C16189T C16192T C16201T T16223C G16230A C16256T T16278C C16311T Brandt 2013
Early Bronze Northern Ireland Glebe, Rathlin Island [Rathlin1] M 2026–1885 BC R1b1a1a2a1a2c1a5

[DF21 was R1b1a2a1a2c1g in 2015]

L21/M529/S145 > DF13/S521 > DF21/S192 U5a1b1e 16519T Cassidy 2015
Nordic LN Sweden Fredriksberg [RISE97] M 2025-1885 BC 580,367



K2a5 146C, 152C, 263G, 324T, 709A, 750G, 1438G, 1811G, 2706G, 3107T, 3480G, 9055A, 9698C, 9716C, 10550G, 11299C, 11467G, 11719A, 12308G, 12372A, 14167T, 14766T, 14798C, 15326G, 15799G, 16224C, 16311C, 16519C Allentoft 2015
Early Bronze Northern Ireland Glebe, Rathlin Island [Rathlin2] M 2024–1741 BC R1b1a1a2a1a2c1a [DF13 was R1b1a2a1a2c1 in 2015] L21/M529/S145 > DF13/S521 U5b2a2 16519T, 16192insC [negative for 4732G, 7521G, 7768G, 16192T] Cassidy 2015
Unetice Germany Quedlinburg [I0164 / QUEVIII 6] F 2023-1894 calBCE (3599±25 BP)

863,754

U5b2a1b T16189C Adler 2012; Haak 2015; Mathieson 2015; Lazaridis 2016
England Hexham Golf Course, Northumberland [I2609 / MOA 1956.46 Box 136] F 2023–1772 calBCE 273312
K2a Olalde 2017
Srubnaya outlier Russia Spiridonovka IV, Samara River, Samara [I0354 / SVP 1] F 2016-1692 BC

385,103 U5a1 Mathieson 2015
Scotland Stenchme, Lop Ness, Sanday, Orkney [I2981 / GENSCOT80] F 2009–1497 calBCE 788384
T2a1b1a Olalde 2017
Vatya Hungary Erd 4 [RISE479] M 2000-1500 BC
884,093 I2a2a1a2a L1229
T2b 73G, 263G, 709A, 750G, 930A, 1438G, 2706G, 3107T, 3157T, 4917G, 8697A, 10463C, 11251G, 11719A, 11812G, 11899C, 13368A, 14233G, 14766T, 14905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16294T, 16296T, 16304C, 16519C Allentoft 2015; Mathieson 2015
Vatya Hungary Erd 4 [RISE483] F 2000-1500 BC


134,337



H2a1 263G, 750G, 951A, 15326G, 16354T Allentoft 2015; Lazaridis 2016
Vatya Hungary Erd 4 [RISE484] F 2000-1500 BCE
168,871

T1a1 73G, 152C, 195C, 263G, 750G, 2706G, 4050T, 4917G, 7829T, 8697A, 9899C, 11251G, 12633A, 14905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16163G, 16186T, 16189C, 16294T, 16519C Allentoft 2015
Russia Bertek-56 on the Ukok Plateau, Altai [child] M 2000-1000 BC K1a24a Pilipenko 2016
Russia Bertek-56 on the Ukok Plateau, Altai [adult] M 2000-1000 BC Q C Pilipenko 2016

Spain Costa Lloguera, Castellon de la Plana [COST1]



H 16293R [in 2008 study] Fernández 2006; Gamba 2008; Oliver 2008

Spain Costa Lloguera, Castellon de la Plana [COST3]



R? 16223Y [in 2008 study], reported as L3a? Fernández 2006; Gamba 2008; Oliver 2008

Spain Costa Lloguera, Castellon de la Plana [COST4, COST5, 3COST4]



H or U 3 samples CRS Fernández 2006; Gamba 2008

Spain Costa Lloguera, Castellon de la Plana [3COST1, 3COST3]



H or RO 2 samples 16362C Fernández 2006

Spain Malpaso cave, Castellnovo (Castellon) [1MP1]



D 16223T, 16325C, 16362C Gamba 2008

Spain Malpaso cave [4MP12]



V 16298C Gamba 2008

Spain Malpaso cave [2MP4]



H20? (reported as L3?) 16218T, 16223T Gamba 2008

Spain Jovellus sepulchre, Benicassim (Castellon) [1JO1]



H20? (reported as L3?) 16218Y, 16223Y Gamba 2008

Spain Assud de Villareal 2nd sepulchre, Almassora (Castellon) [1AV1]



H20? (reported as L3?) 16218Y, 16223Y Gamba 2008

Spain El Portalón, Sierra de Atapuerca M 2000 BC

U5b1b
Daskalaki 2014
Unetice Germany Benzingerode-Heimburg [BZH 10]
1983-1939 BC

X A16129G T16187C G16230A C16311T Brandt 2013
Unetice Germany Benzingerode-Heimburg [BZH 8]
1982-1942 BC

H2a1a3 G73A C146T C152T C195T A247G A769G A825t G951A A1018G G1438A G2706A A2758G C2885T T3594C G4104A T4312C G4769A C6173T T7028C G7146A T7256C A7521G T8468C T8655C G8701A C9540T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C T13095C G13105A G13276A T13506C T13650C T14766C A16129G T16187C C16189T T16223C G16230A A16240t T16278C C16311T C16354T C16519T Brotherton 2013; Brandt 2013
Unetice Germany Esperstedt [ESP 9] M 1981-1878 BC

U5b1a'b A16129G T16187C C16192T T16223C G16230A C16270T T16278C C16311T Adler 2012; Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 31]
1980-1870 BC

T1a T16126C A16129G A16163G C16186T T16187C T16223C G16230A T16278C C16294T C16311T Brandt 2013
Unetice Germany Quedlinburg VII 2 [QLB 39]
1980-1870 BC

W3a1 C146T C152T A189G C194T T204C G207A A247G C146T C152T A189G C194T T204C G207A A247G 309.1C 315.1C A16129G T16187C C16189T G16230A T16278C C16292T C16311T Brandt 2013
Unetice Germany Eulau [EUL 53, 60, 61, 62]
1979-1921 BC for EUL 53

I1a1 4 samples of C146T C152T C195T T199C G203A T204C A247G T250C T16172C T16187C C16189T G16230A T16278C G16391A Brandt 2013
Unetice Germany Eulau [EUL 36 and 37]
1978-1921 BC

H 2 samples G73A C146T C152T C195T A247G A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Unetice Germany Karsdorf [KAR 33]
1978-1915 BC

U5a A16129G T16187C C16189T C16192T C16201T T16223C G16230A C16256T C16270T T16278C C16311T Brandt 2013
Unetice Germany Eulau [EUL 40]
1977-1915 BC

V A16129G T16187C C16189T T16223C G16230A T16278C T16298C C16311T Brandt 2013
Sintashta Russia Bol'shekaraganskii [RISE395] F 1960-1756 BC 888,393

U2e1h 73G, 217C, 228A, 263G, 340T, 508G, 750G, 1438G, 1811G, 2706G, 3107T, 3720G, 6045T, 10876G, 11467G, 11719A, 12308G, 12372A, 13020C, 13734C, 14766T, 15326G, 15907G, 16051G, 16129C, 16183C, 16362C, 16519C Allentoft 2015
England East Kent Access (Phase II), Thanet, Kent [I2601 / EXEKA09_126005] F
1960–1750 calBCE 296184
U5a1b1a Olalde 2017
Unetice Poland Wojkowice [RISE109] F 1954-1772 BC 211,732

U4 73G, 195C, 263G, 310C, 499A, 750G, 1438G, 1811G, 2706G, 3083C, 3107T, 3387C, 6047G, 7028T, 9989C, 11332T, 11467G, 11719A, 12192A, 12308G, 12372A, 12630A, 14364A, 14620T, 14766T, 14851C, 15326G, 15693C, 16356C, 16519C Allentoft 2015
Unetice? Germany Esperstedt [I0115/ESP 3] F 1954-1760 calBCE (3540±31 BP) 365,760

U5a1i A16129G T16187C C16189T C16192T T16223C G16230A C16256T C16270T T16278C C16311T A16399G Adler 2012; Brandt 2013; Haak 2015; Mathieson 2015
Unetice Germany Eulau [EUL 49]
1951-1884 BC

T2b T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T T16304C C16311T Brandt 2013
Sintashta Russia Bulanovo [RISE394] F 1949-1754 BC 336,702

U2e1e 73G, 152C, 217C, 263G, 340T, 508G, 750G, 1438G, 1811G, 2526T,. 2706G, 3107T, 3720G, 6045T, 6872G, 10876G, 11467G, 11719A, 12308G, 12372A, 12618A, 13020C, 13734C, 14180C, 14766T, 15326G, 15907G, 16051G, 16129C, 16183C, 16189C, 16362C, 16519C Allentoft 2015
Bell Beaker Netherlands Oostwoud-Tuithoorn, West Frisia [I4067 / skeleton 127-M1] F 1945–1692 calBCE 28937 R1b1 Olalde 2017
Unetice? Germany Karsdorf [KAR 36]
1945-1882 BC

U5b2b3a1a C16114T A16129G T16187C C16189T T16223C T16224C G16230A C16270T T16278C C16311T Brandt 2013
Unetice? Germany Karsdorf [KAR 37]



K1a24a A16129G G16145A T16187C C16189T T16223C T16224C G16230A T16278C Brandt 2013
England Summerhill, Blaydon, Tyne & Wear [I2610 / MOA 1973.4H Box 167D, 1] F 1936–1746 calBCE 78777

U2e1b Olalde 2017
Food Vessel England Hasting Hill, Sunderland, Tyne and Wear [I2421 / TWCMS Sk2008-3075, find 12] M
1932–1756 calBCE 393274 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS8591:18095336A->C; R1b1a1a2:CTS8665:18137831T->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:M520:4446430T->A; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6432:8194310C->A; R1b1a1a2:PF6505:21784286G->A; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS3063:14637352T->C; R1b1a:L1345:21558298G->T; R1b1a:PF6263:21159055C->A; R1b1:CTS2134:14193384G->A; R1b1:L780:21183643A->G; R1:CTS997:7132713G->A; R1:CTS2908:14556851C->T; R1:CTS3321:14829196C->T; R1:L875:16742224A->G; R1:M306:22750583C->A; R:CTS7880:17723850C->T; etc U5a1a2a Olalde 2017
England East Kent Access (Phase II), Thanet, Kent [I2463 / EXEKA09_246136] F 1930–1740 calBCE 234559
J1c3+189 Olalde 2017
Unetice Poland Szczepankowice [RISE154] F 1925-1765 BC 171,391



K1a4a1 73G, 263G, 497T, 750G, 2706G, 3480G, 5031A, 6260A, 9055A, 10398G, 10550G, 11299C, 11467G, 11485C, 11840T, 12308G, 12372A, 13740C, 14167T, 14180C, 14766T, 14798C, 15326G, 16042A, 16224C, 16311C, 16519C Allentoft 2015
Unetice Germany Benzingerode-Heimburg [BZH 3]
1923-1878 BC

X A16129G T16187C G16230A C16311T Brandt 2013
Srubnaya Russia Spiridonovka IV, Samara River, Samara [I0358 / SVP 6] F 1913-1629 BC 273,856 H6a1a Mathieson 2015
Late Krotovo Russia Tartas-1 [TK10]
1900-1700 BC

A10* 663G, 16223T, 16227C, 16290T, 16311C, 16319A Pilipenko 2015
England East Kent Access (Phase II), Thanet, Kent [I2602 / EXEKA09_136128] M
1900–1690 calBCE 419770 R1b1a1a2a1a2

[P312]


R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:M269:22739367T->C; R1b1a1a2:M520:4446430T->A; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6505:21784286G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5577:16376495A->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6463:16183412C->A; R1b1a:A702:10038192G->A; R1b1a:CTS4244:15510064T->G; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b:M343:2887824C->A; R1:CTS997:7132713G->A; R1:CTS3321:14829196C->T; R1:CTS5611:16394489T->G; R1:L875:16742224A->G; R:CTS7876:17722802G->A; etc. U5b2b Olalde 2017
Maros Hungary Szoreg - C [RISE373] F 1886-1696 BC 259,708



K1a2a 73G, 497T, 750G, 1438G, 1811G, 2706G, 3107T, 3480G, 5773A, 9055A, 9698C, 10398G, 10550G, 11025C, 11299C, 11467G, 11719A, 12308G, 12372A, 14167T, 14766T, 14798C, 15326G, 16224C, 16311C, 16519C Allentoft 2015
Unetice Poland Przeclawice [RISE150] F 1885-1693 BC 654,921 U5a1b1 73G, 263G, 750G, 1438G, 2706G, 3107T, 3197C, 9477A, 9667G, 11467G, 11719A, 12308G, 12372A, 13617C, 14766T, 14793G, 15218G, 15326G, 16192T, 16256T, 16270T, 16291T, 16399G Allentoft 2015
Bell Beaker Netherlands Oostwoud-Tuithoorn, West Frisia [I4071 / skeleton 231 barrow I-M10] F 1883–1665 calBCE 774001 H6a1a Olalde 2017
Bell Beaker Netherlands Oostwoud-Tuithoorn, West Frisia [I4070 / skeleton 230 barrow I-M7] M 1881–1646 calBCE 697834 R1b1a1a2a1a1

[U106]

R1b1a1a2a1a1:M405:8796078C->T; R1b1a1a2a1a:L52:14641193C->T; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:CTS12478:28590278G->A; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:M520:4446430T->A; R1b1a1a2:PF6399:2668456C->T; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6434:8411202A->G; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS7941:17755905G->A; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:CTS11985:23403749G->A; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6463:16183412C->A; R1b1a1a:PF6524:23452965T->C; R1b1a:A702:10038192G->A; R1b1a:CTS3063:14637352T->C; R1b1a:CTS8436:18026855G->A; R1b1a:FGC35:18407611C->T; R1b1a:L754:22889018G->A; R1b1a:L761:16773870A->G; R1b1a:L1345:21558298G->T; R1b1a:PF6263:21159055C->A; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b1:L278:18914441C->T; R1b1:L1349:22722580T->C; R1b:M343:2887824C->A; R1:CTS2565:14366723C->T; etc U5a1b1a Olalde 2017
Maros Hungary Szoreg - C [RISE374] M 1866-1619 BC 146,423 G2a F3088

or (Kendall) G2a2a1a (PF3177)

T2b 73G, 263G, 709A, 750G, 930A, 1438G, 2706G, 4917G, 8292A, 9067G, 10463C, 11251G, 11719A, 11812G, 13368A, 14233G, 14766T, 14905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16304C, 16311C, 16519C Allentoft 2015; Mathieson 2015; add info from Ted Kendall
Srubnaya Russia Novoselki, Northern Forest, Samara [I0232 / SVP 12] M 1850-1200 BC 1,153,002 R1a1a1b2 Z93 U5a1f2 Mathieson 2015
Srubnaya Russia Rozhdestveno I, Samara Steppes, Samara [I0234 / SVP 25] F 1850-1600 BC 467,183 I1a1

Mathieson 2015
Srubnaya Russia Rozhdestveno I, Samara Steppes, Samara [I0235 / SVP 26] F 1850-1600 BC 420,407 K1b2a

Mathieson 2015
Srubnaya Russia Spiridonovka IV, Samara River, Samara [I0259 / SVP 7] F 1850-1200 BC 754,432 U5a2a1 Mathieson 2015
Srubnaya Russia Spiridonovka IV, Samara River, Samara [I0260 / SVP 8] M 1850-1200 BC 58,414 R1a1 M459 U5a1 Mathieson 2015
Srubnaya Russia Spiridonovka IV, Samara River, Samara [I0261 / SVP 9] M 1850-1200 BC 415,327 R1a1a L168

H5b Mathieson 2015
Srubnaya Russia Barinovka I, Samara River, Samara [I0422 / SVP 30] F 1850-1200 BC 750,562 T1a1 Mathieson 2015
Srubnaya Russia Barinovka I, Samara River, Samara [I0423 / SVP 31] M 1850-1200 BC 418,611 R1a1a1b2 Z93 J2b1a2a Mathieson 2015
Srubnaya Russia Uvarovka I, Samara River, Samara [I0424 / SVP 32] M 1850-1600 BC 16,569,296 R1a1a1b2 Z93 T2b4 Mathieson 2015
Srubnaya Russia Spiridonovka II, Samara River, Samara [I0430 / SVP 39] M 1850-1600 BC 22,835,907 R1a1a1b2a2a Z2123 H3g 1st degree relative of I0421 Mathieson 2015
Srubnaya Russia Spiridonovka II, Samara River, Samara [I0431 / SVP 40] F 1850-1600 BC 107,284 H2b Mathieson 2015; Lazaridis 2016
Srubnaya Russia Spiridonovka II, Samara River, Samara [I0421 / SVP 29] F 1850-1600 BC 52,004 H3g 1st degree relative of I0430. Mathieson 2015
Unetice Germany Benzingerode-Heimburg [BZH 13]
1804-1772 BC

T2b T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T C16296T T16304C C16311T Brandt 2013
Andronovo Russia Tatarka cemetery, burial 64 Charypovsky region [S07] Brown eyes; dark brown hair; fair or medium skin M 1800-1400 BC C (not C3) Ci in table 2 U4 16356C Keyser 2009
Andronovo Russia Tatarka, burial 55 [S08] Brown eyes F 1800-1400 BC

Z1
Keyser 2009
Andronovo Russia Solenoozernaďa IV, kourgane I, burial 3, Krasnoyarsk region [S09] Blue or green eyes, blond or light brown hair, fair or medium skin M 1800-1400 BC

T1 16126C, 16163G, 16186T, 16189C, 16294T Keyser 2009
Andronovo Russia Solenoozernaďa IV, kourgane I, burial 4 [S10] Brown hair M 1800-1400 BC R1a1a M17; Rii in table 2 U2e 16051G, 16092C, 16129C, 16183C, 16189C, 16362C Keyser 2009
Andronovo Russia Solenoozernaďa I, burial 4 [S11] Blue or green eyes F 1800-1400 BC

T2a1b1 16126C, 16294T 16324C, reported as T4. Keyser 2009
Andronovo Russia Solenoozernaďa IV, kurgane I, burial 4 [S13] ? 1800-1400 BC

H6 16187T, 16362C Keyser 2009
Andronovo Russia Solenoozernaďa I, burial 4 Krasnoyarsk region [S14] Brown eyes, brown hair, fair or medium skin F 1800-1400 BC

U4 16356C Keyser 2009
Andronovo Russia Solenoozernaďa I, burial 29 [S15] Blue or green eyes ? 1800-1400 BC

K2b 16093C, 16224C, 16311C, 16319A Keyser 2009
Andronovo Russia Oust-Abakansty, chief kurgan, Khakassia republic [S16] Blue or green eyes, blond or light brown hair, fair or medium skin M 1800-1400 BC R1a1a M17; Rii in table 2 U5a1 16192T, 16256T, 16270T Keyser 2009
Andronovo (Fedorovo) Russia Tartas-1 [TK17]
1800-1300 BC

A10* 663G, 16223T, 16227C, 16278T, 16290T, 16311C, 16319A Pilipenko 2015
Andronovo Russia Kytmanovo [RISE500] F
904483



U4d1 73G, 195C, 263G, 499A, 629C, 750G, 1438G, 1533T, 1700C, 1811G, 2706G, 2772T, 3107C, 11031A, 11332T, 11467G, 11719A, 12308G, 12372A, 14620T, 14766T, 15326G, 15693C, 16356C, 16519C Allentoft 2015
Unetice Germany Esperstedt [ESP 18] M 1776-1664 BC

T2b T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T C16296T T16304C C16311T Adler 2012; Brandt 2013
Gáta/Wieslburg Hungary M85 Enese elkerül 02. Kóny, Proletár II [KON 6]
1770-1760 BC R1b1a2 M269 U5b1 16093c 16189c 16270t Szécsényi-Nagy 2015 thesis
Bulgaria Kairyaka Necropolis, Merichleri [I2163 / Merich2, individual N5] M 1750-1625 calBCE 825494 R1a1a1b2

[Z93]

R1a1a1b2:F992:7552356G->A; R1a1a1b:S224:8245045C->T; R1a1a1b:S441:7683058G->A; R1a1a1:Page7:14498990C->T; R1a1a:L168:16202177A->G; R1a1a:M515:14054623T->A; R1a:L62:17891241A->G; R1a:L63:18162834T->C; R1a:L145:14138745C->A; R1:CTS997:7132713G->A; R1:CTS2565:14366723C->T; R1:CTS3123:14674176A->C; R1:CTS5611:16394489T->G; R1:L875:16742224A->G; R1:P225:15590342G->T; R1:P231:9989615A->G; R1:P236:17782178C->G; R1:P238:7771131G->A; R1:P286:17716251C->T; R1:P294:7570822G->C; R:CTS2913:14561760A->G; etc. U5a2 Mathieson 2017

Spain El Portalón, Sierra de Atapuerca [ATP9] F 1750-1618 calBCE (3390±30, Beta-386395) 449,162

U5b1b 146T, 150T, 152T, 195T, 247G, 769G, 825T, 1018G, 2758G,
2885T, 3197C, 3594C, 4104A, 4312C, 5656G, 7146A, 7256C, 7521G, 7768G, 8468C, 8655C, 8701A, 9477A, 9540T, 10398A, 10664C, 10688G, 10810T, 10873T, 10915T, 11467G, 11914G, 12308G, 12372A, 12705C, 13105A, 13276A, 13506C, 13617C, 13650C, 14182C, 16129G, 16187C, 16192T, 16223C, 16230A,
16270T, 16278C, 16311T, 8838A, 10172A, 11878C,
12630A, 13722G, 16221T,
16362C, 16519T
Günther 2015
Bronze Age England Boscombe Airfield, Wiltshire [I2464 / 62260_39] M
1750–1610 calBCE 52649 R1b1a1a2a1

[L51]


R1b1a1a2a1:L51:8502236G->A; R1b1a:L761:16773870A->G; R1b:M343:2887824C->A; R:F459:18017528G->T; R:M734:18066156C->T I2 Olalde 2017
Andronovo Russia Kytmanovo [RISE505] F 1746-1626 BC 1,170,027



U4a1b 152C, 195C, 263G, 499A, 750G, 1438G, 1811G, 2706G, 3107C, 3204T, 11332T, 11467G, 11719A, 12308G, 12372A, 12937G, 14620T, 14766T, 15326G, 15693C, 16134T, 16356C, 16519C Allentoft 2015
Vatya Hungary Szazhalombatta-Foldvar [RISE247] M 1746-1611 BC 237,322

I2a2a1

CTS9183 H11a 750G, 961G, 1438G, 13759A, 15326G, 16293G, 16311C Allentoft 2015; Mathieson 2015
Early Bronze Northern Ireland Glebe, Rathlin Island [Rathlin3] M 1736–1534 BC R1b1a1a2a1a2c1

[L21 was R1b1a2a1a2c in 2015]

L21/M529/S145 J2b1a 14819C, 16519T, [negative for 4216C, 5633T, 7146A, 7476T, 7521G] Cassidy 2015
Andronovo Russia Kytmanovo [RISE503] F 1727-1511 BC 678,907

U2e2 73G, 152C, 217C, 263G, 508G, 1230T, 1438G, 1811G, 2706G, 3107C, 3720G, 5390G, 6896C, 8473C, 10876G, 11467G, 11719A, 12308G, 12372A, 12557T, 13020C, 13734C, 14753T, 14766T, 15326G, 15907G, 16051G, 16092C, 16129C, 16311C, 16362C, 16519C, 16527T Allentoft 2015
Croatia Veliki Vanik [I4331 / VV1] M 1700-1500 BCE 725669 J2b2a J2b2a:L283:6932663C->G; J2b:L282:5359116C->T; J2b:M314:22753079A->C; J:CTS687:6953311A->T; J:CTS852:7048870G->A; J:CTS4204:15476324G->A; etc I1a1 Mathieson 2017
Vatya Hungary Erd 4 [RISE480] F 1700-1500 BC
158,897

U5a2a 73G, 263G, 2706G, 3197C, 5379T, 11467G, 12308G, 12372A, 12969T, 13617C, 15326G, 16114A, 16192T, 16256T, 16270T, 16526A Allentoft 2015
Wales Great Orme Mines, Llandudno [I1775 / GOM245] M 1693–1600 calBCE 410642 R1b1a1a2a1a2

[P312]

R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS623:6912992T->G; R1b1a1a2:CTS8665:18137831T->C; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6399:2668456C->T; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6482:18381735A->G; R1b1a1a2:PF6494:20811307G->A; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a:A702:10038192G->A; R1b1a:L754:22889018G->A; R1b1:L278:18914441C->T; R1b1:L822:7960019G->A; R1b1:L1349:22722580T->C; R1:CTS997:7132713G->A; R1:CTS2565:14366723C->T; R1:L875:16742224A->G; R:CTS3622:15078469C->G; etc. I3 Olalde 2017
Unetice Germany Karsdorf [KAR 39]
1653-1627 BC

H G73A C146T C152T C195T A247G 309.1C 315.1C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013
Unetice Germany Benzingerode-Heimburg [BZH 9]
1653-1627 BC

K A16129G T16187C C16189T T16223C T16224C G16230A T16278C Brandt 2013
Unetice? Germany Benzingerode-Heimburg [BZH 17]
1653-1627 BC

U5a1a'g A16129G T16187C T16223C G16230A C16256T C16270T T16278C C16311T A16399G Brandt 2013
Unetice Germany Benzingerode-Heimburg [BZH 18]
1653-1627 BC

I3a T16086C T16187C C16189T G16230A T16278C C16311T G16391A Brandt 2013
Nuragic Italy Su Grutta is Bittuleris, Sardinia
1624 BC

H1aw1 G3010A, A8701G!, C15912T Brotherton 2013
England Amesbury (Boscombe) Down, Wiltshire [I2639 / 25217] "Amber boy" in fact is Amber girl F
1600–1430 BC 579584

T2e1a Olalde 2017
Mezhovskava Russia Kapova cave [RISE523] F 1598-1398 BC 14794


M12'G 73G, 152C, 263G, 489C, 709A, 750G, 1438G, 2706G, 3107C, 8547C, 9377G, 9540C, 10398G, 10400T, 11719A, 12705T, 14200C, 14569A, 14587G, 14621T, 14766T, 14783C, 15043A, 15301A, 15326G, 16223T, 16278T,16362C, 16519C Allentoft 2015
Mezhovskava Russia Kapova cave [RISE524] M
R1b R1b1a2 (PF6494) J2b1a 73G, 150T, 152C, 263G, 295T, 489C, 750G, 1438G, 2706G, 3107T, 7028T, 9055A, 10172A, 10398G, 11251G, 11719A, 12612G, 13708A, 14766T, 15257A, 15326G, 15452A, 15812A, 16069T, 16126C, 16193T, 16278T Allentoft 2015; Y-DNA personal communication from author + additional info from Sergey Malychev and Felix Emmanuel
Mezhovskava Russia Kapova cave [RISE525] M
R1a
I5c 73G, 152C, 250C, 263G, 750G, 1438G, 1719A, 2706G, 10034C, 10238C, 10398G, 12501A, 12705T, 13780G, 14766T, 15043A, 15326G, 15924G, 16086C, 16129A, 16169T, 16223T, 16258G, 16391A, 16519C Allentoft 2015; Y-DNA personal communication from author

Spain Montanissell Cave, Sant Joan de Montanissell, Lleida, Catalonia [E10]
1546 ± 52 BC

U5a1c 16192T; 16256T; 16270T; 16320T; 16399G Simón 2011
Karasuk Russia Sabinka 2 [RISE493] M 1531-1427 BC Q1a
C4a1c 195C, 263G, 489C, 750G, 1438G, 1715T, 2706G, 3107C, 3552A, 10398G, 10400T, 11719A, 11914A, 11969A, 12672G, 12705T, 13263G, 14318C, 14766T, 14783C, 15043A, 15204C, 15301A, 15326G, 15487T, 15607G, 15968C, 16093C, 16129A, 16209C, 16223T, 16298C, 16327T, 16519C Allentoft 2015; Y-DNA personal communication from author

Spain Montanissell Cave [E9] F 1524 ± 56 BC

J/J1c 16069T; 16126C Simón 2011
Babino Ukraine Liubasha, kurgan 2 [L11] 1523 ± 76 BC HV1 7028T, 16067T, 16192T Nikitin 2017

Spain Montanissell Cave [E5] F 1510-1420 BC

J/J1c 16069T; 16126C Simón 2011
Scotland Evergreen House Longniddry, East Lothian [I2573 / Skeleton 4 / GENSCOT21] F 1501–1302 calBCE 611500 H3+16311 Olalde 2017

Ireland Glencurran Cave
1500 BC

U5


Altaď Republic Bertek-56, Ukok Plateau M 1500 BC

H? CRS Chikisheva 2007
Mycenaen Greece Grave Circle B, Mycenae [55, 58] 1500 BC

K Brother and sister?
16224, 16311
Bouwman 2008
Mycenaen Greece Grave Circle B, Mycenae [Z59]
1500 BC

U5a1/U5a1a 16256, 16266, 16270 Bouwman 2008
Mycenaen Greece Grave Circle B, Mycenae [A62]
1500 BC

H? CRS Bouwman 2008
Karakol Altaď Republic mound 3, Karakol-1, Gorny Altai M 1500 BC

U5 16192, 16256, 16270 Chikisheva 2007
Karakol Altaď Republic mound 3, Karakol, Gorny Altai M 1500 BC

H 16304 Chikisheva 2007

Spain Montanissell Cave [E2] M 1500-1420 BC

U CRS, identified as U on coding region Simón 2011

Spain Montanissell Cave [E7] M 1500-1410 BC

J CRS, partial result, identified as J on coding region Simón 2011

Spain Montanissell Cave [E8] F 1500-1410 BC

V 16298C Simón 2011

Spain Montanissell Cave [E1] F 1500-1410 BC

K 16224C; 16311 Simón 2011
England Canada Farm, Sixpenny Handley, Dorset [I3082 / Burial F4] M 1500–1390 calBCE 582197 R1b1a1a2a1a2c


R1b1a1a2a1a2c:L21:15654428C->G; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L151:16492547C->T; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2a:L23:6753511G->A; R1b1a1a2:CTS8728:18167403C->T; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:L150.1:10008791C->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:L773:7220727A->G; R1b1a1a2:PF6425:7762947T->C; R1b1a1a2:PF6430:8070532T->A; R1b1a1a2:PF6438:9464078C->T; R1b1a1a2:PF6475:17986687C->A; R1b1a1a2:PF6494:20811307G->A; R1b1a1a2:PF6495:20828795G->A; R1b1a1a2:PF6497:21222868C->G; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6451:14116584T->A; R1b1a1a:PF6463:16183412C->A; R1b1a:FGC36:13822833G->T; R1b1a:FGC41:7900883C->A; R1b1a:PF6249:8214827C->T; R1b1a:PF6263:21159055C->A; R1b1:CTS2134:14193384G->A; R1b1:L780:21183643A->G; R1b1:L822:7960019G->A; R1b1:M415:9170545C->A; R1:CTS997:7132713G->A; R1:F93:7671535C->T; R1:P225:15590342G->T; R1:P238:7771131G->A; R1:P294:7570822G->C; R:CTS207:2810583A->G; etc. H1an1a 152C, 263G, 750G, 1438G, 3010A, 4454C, 4769G, 8860G, 13050G, 15326G, 15394C, 16519C Olalde 2017
Scotland Evergreen House Longniddry, East Lothian [I2653 / Skeleton 2 / GENSCOT22] M 1500–1300 BCE 574585 R1b1a1a2a1a2c1a4b2c1a on ISOGG 12 May 2017, reported in Olalde as R1b1a1a2a1a2c1e2b3a1 (ISOGG 2016] [L21 >>>> Z2186] R1b1a1a2a1a2c1e2b3a1:CTS1202.1:7265766A->C; R1b1a1a2a1a2c1e2b3:S883:15339857A->G; R1b1a1a2a1a2c1e2b:S868:19033817T->C; R1b1a1a2a1a2c:L21:15654428C->G; R1b1a1a2a1a2:P312:22157311C->A; R1b1a1a2a1a:L11:17844018T->C; R1b1a1a2a1a:P310:18907236A->C; R1b1a1a2a1a:P311:18248698A->G; R1b1a1a2a1:L51:8502236G->A; R1b1a1a2:CTS2664:14416216G->A; R1b1a1a2:CTS10834:22796697T->C; R1b1a1a2:CTS11468:23124367G->T; R1b1a1a2:L265:8149348A->G; R1b1a1a2:PF6399:2668456C->T; R1b1a1a2:PF6438:9464078C->T; R1b1a1a:CTS3876:15239181G->C; R1b1a1a:CTS5082:16005138A->C; R1b1a1a:CTS7904:17732408T->C; R1b1a1a:CTS9018:18617596C->T; R1b1a1a:FGC57:7759944G->A; R1b1a1a:L502:19020340G->C; R1b1a1a:PF6463:16183412C->A; R1b1a1a:PF6498:21312064C->A; R1b1a:A702:10038192G->A; R1b1a:FGC36:13822833G->T; R1b1a:L754:22889018G->A; R1b1a:L1345:21558298G->T; R1b1a:PF6271:23984056G->A; R1b1:CTS2134:14193384G->A; R1b1:CTS2229:14226692T->A; R1b:M343:2887824C->A; R1:CTS2565:14366723C->T; R1:F93:7671535C->T; R1:L875:16742224A->G; etc. H2a1a Olalde 2017
Scotland Evergreen House Longniddry, East Lothian [I2654 / Skeleton 2 / GENSCOT23] F 1500–1300 BCE 518165 U5b3 Olalde 2017

Denmark Bredtoftegĺrd [Bt1]
1500-1300 BC

U4 7028T, 12308G, 16179T, 16356 Melchior 2010
Croatia Jazinka Cave [I3313 / JAZ1]

F 1500-900 BCE 796183 HV0e Mathieson 2017
Nordic Bronze Age Denmark Sebber skole [RISE47] M 1499-1324 BC 103,513

R1b1a2 M520+, PF6438+

[*Tilroe adds: xP312, xA2150]

I 73G, 750G, 1438G, 2706G, 10034C, 10238C, 11719A, 13780G, 15326G, 16223T, 16391A Allentoft 2015; Mathieson 2015; *additional information from Vince Tilroe
Karasuk Russia Bystrovka [RISE502] F 1496-1306 BC

U5a1d 73G, 263G, 750G, 1438G, 2706G, 3027C, 3107T, 3197C, 9477A, 11467, G11719A, 12308G, 12372A, 13617C, 14053G, 14766T, 14793G, 15218G, 15326G, 16192T, 16256T, 16270T, 16399G Allentoft 2015
Nordic Bronze Age Sweden Angmollan [RISE207] M 1493-1302 BC I1
J1c8a1 73G, 750G, 1438G, 2706G, 3010A, 11251G, 13708A, 14798C, 15326G, 15452A, 16069T, 16265G Allentoft 2015 ; Y-DNA personal communication from author

Spain Urtiaga, Gipuzkoa [URT-1]
1475-1430 BC

H? rCRS in HVRI, 73A, reported as H Hervella 2012

Spain Urtiaga, Gipuzkoa [URT-2]
1475-1430 BC

U5a 73G, 16192, 16270 Hervella 2012

Spain Urratxa, Bizkaia
1475-1405 BC

H 2 samples Izagirre and de la Rúa 1999

Spain Urratxa, Bizkaia
1475-1405 BC

J 2 samples Izagirre and de la Rúa 1999

Spain Urratxa, Bizkaia
1475-1405 BC

U 2 samples Izagirre and de la Rúa 1999
Andronovo Russia Kytmanovo [RISE512] M 1446-1298 BC R1a1a1b

[*R1a1a1b2a2]

S224 (Mathieson)

[*Tagankin reads Z2121+]

U2e1 73G, 152C, 340T, 508G, 1438G, 1811G, 2706G, 3720G, 4255T, 5390G, 10876G, 11467G, 11719A, 12308G, 12372A, 13020C, 13734C, 14766T, 15326G, 15907G, 16051G, 16129C, 16189C, 16362C, 16381C, 16391A, 16519C Allentoft 2015; Mathieson 2015; Y-DNA *additional info from Vladimir Tagankin
Scotland Pabay Mor, Isle of Lewis, Western Isles [I2655 / GENSCOT24] M 1442–1273 calBCE 835188 I2a2a1a1a1 I2a2a1a1a1:L126:14901633C->T; I2a2a1a1a1:L137:9791250G->A; I2a2a1a1a1:L369:14850314T->C; I2a2a1a1a:L1195:18865320G->A; I2a2a1:CTS9183:18732197A->G; I2a2a:P221:8353707C->A; I2a2a:P223:16699334C->G; I2a2:L35:22725379C->A; I2a2:L37:17516123T->C; I2a2:L181:19077754G->T; I2a2:L368:6931594C->T; I2a2:P216:13992338C->G; I2a2:P217:7628484C->T; I2a2:P218:17493630T->G; I2:L68:18700150C->T; I2:PF3664:8567995G->A; I:CTS674:6943522C->T; I:CTS1800:14073053G->A; I:CTS2387:14286853T->C; etc. H6a1b2 Olalde 2017

Spain Montanissell Cave [E11] F 1440-1390 BC

J/J1c 16069T; 16126C Simón 2011
Nordic Bronze Age Sweden Angmollan [RISE210] M 1432-1292 BC 51,095

I
CTS4209, CTS7469, PF3817


T2a1a 73G, 263G, 709A, 750G, 1438G, 2706G, 2850C, 3107T, 4917G, 6000T, 7022C, 8697A, 10463C, 11251G, 11719A, 11812G, 13965C, 14233G, 14687G, 14766T, 14905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16294T, 16296T, 16519C Allentoft 2015; Mathieson 2015
Nuragic Italy Carbonia, Sardinia [CA02]
1430 BC

H CRS Caramelli 2007; Ghirotto 2010
Nuragic Italy Carbonia, Sardinia [CA14];
1430 BC

U2 16051G Caramelli 2007; Ghirotto 2010
Nordic Bronze Age Denmark Karlstrup [RISE21] M 1426-1281 BC

N1a1a1a2 5315G, 8485A, 10238C, 11719A, 12498T, 12501A, 13780G, 14766T, 15043A, 5315G, 8485A, 10238C, 11719A, 12498T, 12501A, 13780G, 14766T, 15043A, 15299C, 15326G, 16086C, 16223T, 16248T, 16320T, 16355T, 16519C Allentoft 2015
Karasuk Russia Sabinka 2 [RISE494] M 1416-1268 BC R1a [*R1a1a1b2a2a] [*Z2123+, and downstream SNPs Y934+ YP520+] I4a1 73G, 199C, 204C, 250C, 1438G, 1719A, 2706G, 3107T, 8519A, 10034C, 10238C, 10398G, 10819G, 11719A, 12501A, 12705T, 13780G, 14766T, 14821T, 15043A, 15326G. 15924G, 16129A, 16223T, 16304C, 16391A, 16519C Allentoft 2015; Y-DNA personal communication from author and *additional info from Vladimir Tagankin
Wales North Face Cave, Llandudno [I2574 / NFC07151] F 1415–1228 calBCE 77807 U5a1a2b Olalde 2017
Karasuk Russia Arban I [RISE495] M
R1a
D4j1 73G, 263G, 489C, 750G, 1438G, 2706G, 3010A, 3107T, 4263G, 4883T, 5262A, 7028T, 8414T, 9540C, 9948A, 10398G, 10400T, 10873C, 11696A, 11719A, 12705T, 14130T, 14668T, 14766T, 14769G, 14783C, 15043A, 15301A, 15326G, 16223T, 16260T, 16362C Allentoft 2015; Y-DNA personal communication from author
Karasuk Russia Arban I [RISE496] F 1414-1261 BC

U5a1a2a 73G, 263G, 750G, 1438G, 1700C, 2706G, 3107T, 3197C, 3565G, 5319G, 6629G, 6719C, 9249T, 9477A, 11467G, 11719A, 12308G, 12346T, 12372A, 13617C, 14766T, 14793G, 15218G, 15326G,16256T, 16270T, 16309G, 16519C Allentoft 2015
Karasuk Russia Arban I [RISE497] F


A+152+16362 73G, 150T, 152C, 235G, 263G, 663G, 750G, 1438G, 1736G, 2706G, 3107T, 3213G, 4248C, 4824G, 7028T, 8794T, 11719A, 12705T, 14766T, 15326G, 16189C, 16223T, 16290T, 16319A, 16362C Allentoft 2015
Karasuk Russia Bystrovka [RISE499] F


H5a1 263G, 456T, 750G, 1438G, 3107C, 4336C, 10634G, 15326G, 15833T, 16304C Allentoft 2015

Kazakhstan Oi-Zhaylau-III, Talapty-II
1400-1000 BC

T1 16126, 16163, 16186, 16189, 16294 Lalueza-Fox 2004

Kazakhstan Oi-Zhaylau-III, Talapty-II
1400-1000 BC

T1 16093, 16126, 16163, 16186, 16189, 16294 Lalueza-Fox 2004

Kazakhstan Vodokhranilische (Kourgan 4)
1400-1000 BC

HV 16129 Lalueza-Fox 2004

Kazakhstan Vodokhranilische (Kourgan 4)
1400-1000 BC

I 16129, 16223, 16294 Lalueza-Fox 2004

Kazakhstan Vodokhranilische (Kourgan 4)
1400-1000 BC

T* 16126, 16288, 16294, 16296 Lalueza-Fox 2004

Kazakhstan Ak-Mustafa (Kourgan 2)
1400-1000 BC

U5 16187, 16192, 16260, 16270 Lalueza-Fox 2004

Kazakhstan Izmaylovka (Kourgan 7,10)
1400-1000 BC

H CRS Lalueza-Fox 2004

Kazakhstan Izmaylovka (Kourgan 7,10)
1400-1000 BC

U5a1 16192, 16256, 16270, 16311 Lalueza-Fox 2004
Karasuk Russia Oust-Abakansty, kurgan IV, burial 1, Khakassia republic [S18]
Blue or green eyes
F 1400-800 BC

U5a1 16114A, 16256T, 16270T, 16294T Keyser 2009
Karasuk Russia Bogratsky, burial I
Khakassia republic [S19]
Blue or green eyes
F 1400-800 BC

U4* 16356C, 16362C Keyser 2009
Pakhomovo Russia Stary Sad [Sts6]
1400-700 BC

A10* 663G, 16093C, 16223T, 16227C, 16290T, 16311C, 16319A Pilipenko 2015
Pakhomovo Russia Stary Sad [Sts11]
1400-700 BC

A10* 663G, 16148T, 16223T, 16227C, 16290T, 16311C, 16319A Pilipenko 2015
Nordic Bronze Age Sweden Abekas I [RISE175] M 1395-1132 BC 112,552

I CTS1301, CTS7502, CTS10941
T1a1 73G, 152C, 263G, 709A, 750G, 1204T, 1209T, 1438G, 2706G, 4917G, 8697A, 9899C, 11251G, 11719A, 12633A, 14766T, 14905A, 15326G, 15452A, 15607G, 16126C, 16163G, 16186T, 16189C, 16294T, 16519C Allentoft 2015; Mathieson 2015

Mongolia Tsagaan Asga [TA4]
Black/dark brown hair; brown eyes
M 1371 BC R1a1a1b2 (Z93)
T1a 16093C, 16126C, 16163G, 16186T, 16189C, 16294T Hollard 2014

Mongolia Tsagaan Asga [TA8]
M
Q1a2a1 (L54)
U4 16356C Hollard 2014

Mongolia Tsagaan Asga [TA9]
Black/dark brown hair; brown eyes
F


D5a2 16086C, 16092C, 16164G, 16182C, 16183C, 16185T, 16189C, 16193del, 16223T, 16266T, 16362C Hollard 2014

Mongolia Tsagaan Asga [TA10]
Dark brown hair; brown eyes
M
R1a1a1b2 (Z93)
)C4a1 16093C, 16129A, 16223T, 16260T, 16263C, 16298C, 16327T Hollard 2014

Mongolia Tsagaan Asga [TA11]
Black/dark brown hair; brown eyes
M
Q1a2a1 (L54)
D 16223T, 16362C Hollard 2014

Mongolia Tsagaan Asga [TA12]
Black/dark brown hair; brown eyes
M
Q1a2a1

(L54)


H7e* 16305G, 16311C Hollard 2014

Mongolia Tsagaan Asga [TA14]
Brown hair; blue eyes
M
R1a1a1b2 (Z93)
D2 16092C, 16129A, 16223T, 16362C Hollard 2014

Mongolia Takhilgat Uzuur-5 [TU15]
Dark brown hair; brown eyes
F 1358 BC

H1b 16183C, 16189C, 16356C Hollard 2014
Nuragic Italy Seulo, Sardinia [SE81, SE 84]
1300 BC

H 2 samples CRS Caramelli 2007; Ghirotto 2010
Nuragic Italy Seulo, Sardinia [SE13]
1300 BC

U2 16051G Caramelli 2007; Ghirotto 2010
Nuragic Italy Seulo, Sardinia [SE01]
1300 BC

H 16126C Caramelli 2007; Ghirotto 2010
Nuragic Italy Seulo, Sardinia [SE02]
1300 BC

J 16069T, 16126C, 16192T Caramelli 2007; Ghirotto 2010
Nuragic Italy Seulo, Sardinia [SE60]
1300 BC

H 16278T Caramelli 2007; Ghirotto 2010
Nuragic Italy S Teresa di Gallura, Sardinia [SR38, ST47]
1300-1200 BC

H 2 samples CRS Caramelli 2007; Ghirotto 2010
Nuragic Italy

S Teresa di Gallura, Sardinia [ST15, ST54]


1300-1200 BC

H or U 2 samples of 16129C Caramelli 2007; Ghirotto 2010
Nuragic Italy S Teresa di Gallura, Sardinia [ST08]
1300-1200 BC

? 16223T Caramelli 2007; Ghirotto 2010
Nuragic Italy S Teresa di Gallura, Sardinia [ST30]
1300-1200 BC

H 16126C Caramelli 2007; Ghirotto 2010
Nuragic Italy S Teresa di Gallura, Sardinia [ST16]
1300-1200 BC

H 16311C Caramelli 2007; Ghirotto 2010
Nuragic Italy Fluminimaggiore, Sardinia [FL04]
1265 BC

H or U 16129C Caramelli 2007; Ghirotto 2010

Scotland, UK Cladh Hallan, South Uist (tooth composite skeleton) F 1300-1100 BC

U5 C16270T Hanna 2012; Kerri Brown

Scotland, UK Cladh Hallan, South Uist (femur composite skeleton) F 1300-1100 BC

T1a C16186T, T16189C Hanna 2012; Kerri Brown

Scotland, UK Cladh Hallan, South Uist (humerus composite skeleton) F 1300-1100 BC

H? CRS Hanna 2012; Kerri Brown
Kyjatice Hungary Ludas-Varju-dulo [I1504/BR 2] M 1270-1110 BC J2a1 L26. J2a1b (M67 > Z1847> Z7671> CTS900> Y11202> Z30695> Z30685) . Y-Full id: BR2
K1a1a
Gamba 2014; Mathieson 2015; additional info from Chris Rоttеnѕtеіnеr
Urnfield Germany Halberstadt [I0099 / HAL 36] M 1193-979 calBCE (2889±30 BP)

985,957 R1a1a1b1a2

(Z280)

S204+, S198+, PF6217+ H23 G73A C146T C152T C195T A247GA769G A825t A1018G G2706A A2758G C2885T T3594C G4104A T4312C T7028C G7146A T7256C A7521G T8468C T8655C G8701A C9540T C10211T G10398A T10664C A10688G C10810T C10873T C10915T A11719G A11914G T12705C G13105A G13276A T13506C T13650C T14766C A16129G T16187C C16189T T16223C G16230A T16278C C16311T Brandt 2013; Haak 2015; Mathieson 2015; Lazaridis 2016

Ukraine Verteba Cave [V6]
1013-544 BC

H 14766 Mse I?, 7025 Alu I?, 16070 Nikitin 2011

Mongolia Takhilgat Uzuur-5 [TU34]
Dark blond/ brown hair; brown eyes
M 1011 BC R1a1a1b2

(Z93)


H/U CRS Hollard 2014
Baltic BA Lithuania Turlojiškė II [Turlojiske3] M 1010-800 cal BCE 471779 R1a1a1b H4a1a1a3 Mittnik 2017
Late BA Russia Afontova Gora [RISE554] M 1005-844 BC

F1b1b 73G, 152C, 204C, 263G, 1438G, 2706G, 3107T, 3970T, 4732G, 4776A, 6392C, 6962A, 8772C, 10310A, 10609C, 10976T, 11719A, 12406A, 12633T, 12882T, 13928C, 14476A, 14766T, 15326G, 15924G, 16172C, 16183C, 16189C, 16232A, 16249C, 16304C, 16311C Allentoft 2015
Urnfield Germany Lichtenstein Cave, near Dorste, Lower Saxony [M1, M2, M7] M 1000 BC I2a2b Ii in table 2; Z5REF in Ysearch H 3 samples Schilz 2006
Urnfield Germany Lichtenstein Cave [M3, M6] M 1000 BC I2a2b Iii in table 2 H 2 samples Schilz 2006
Urnfield
Germany Lichtenstein Cave [M14] M 1000 BC I2a2b? Iii? in table 2 H
Schilz 2006
Urnfield Germany Lichtenstein Cave [M4, M5, M19] M 1000 BC I2a2b Iiii in table 2 H 3 samples Schilz 2006
Urnfield Germany Lichtenstein Cave [M8, M16] M 1000 BC I2a2b Ii in table 2; Z5REF in Ysearch U5b 2 samples Schilz 2006
Urnfield
Germany Lichtenstein Cave [M10] M 1000 BC R1a1? Ri in table 2 T2?
Schilz 2006
Urnfield Germany Lichtenstein Cave [M11] M 1000 BC R1a1? Ri? in table 2 U
Schilz 2006
Urnfield Germany Lichtenstein Cave [M9] M 1000 BC R1b Rbi in table 2 H
Schilz 2006
Urnfield Germany Lichtenstein Cave [F1, F12, F13, F19, F20, F21] F 1000 BC

H 6 samples Schilz 2006
Urnfield Germany Lichtenstein Cave [F2, F3, M12, M15] F 1000 BC

J* 4 samples Schilz 2006
Urnfield Germany Lichtenstein Cave [F14] F 1000 BC

J1b1
Schilz 2006
Urnfield Germany Lichtenstein Cave [F5, F6, F10, F16] F 1000 BC

T2 4 samples Schilz 2006
Urnfield Germany Lichtenstein Cave [F9, F18] F 1000 BC

U 2 samples Schilz 2006
Urnfield Germany Lichtenstein Cave [F15] F 1000 BC

U (2?)
Schilz 2006
Urnfield Germany Lichtenstein Cave [F4, F7, F8] F 1000 BC

U5b 3 samples Schilz 2006

Spain Macanet cave, Llucena (Castellon) [1MN1]



H CRS Gamba 2008

Ukraine Verteba Cave [F3]
969-418 BC

U5a1a 14766 Mse I+, 12308 Hinf I+, 7025 Alu I+, 16256, 16270 Nikitin 2011

Mongolia Takhilgat Uzuur-5 [TU18] ? 944 BC

D 16223T, 16311C, 16362C Hollard 2014
Nuragic Italy Perdasdefogu, Sardinia [PE11, PE20, PE23]
930 BC

H 3 samples CRS Caramelli 2007; Ghirotto 2010
Nuragic Italy Perdasdefogu, Sardinia [PE15]
930 BC

V 16129C, 16298C Caramelli 2007 Ghirotto 2010
Nuragic Italy Perdasdefogu, Sardinia [PE25]
930 BC

H? 16223T Caramelli 2007 Ghirotto 2010
Late BA Montenegro Velika Gruda [RISE595] F


U5a2 73G, 263G, 1838T, 2706G, 3197C, 10152A, 11467G, 11983T, 12308G, 12372A, 13617C, 15326G, 16196A, 16256T, 16526A Allentoft 2015
Late BA Russia Afontova Gora [RISE553] M 926-815 BC R1a
F1b 73G, 152C, 204C, 263G, 750G, 2706G, 6392C, 6962A, 10227C, 10310A, 10609C, 10976T, 12406A, 12882T, 13928C, 14476A, 14766T, 15326G, 16172C, 16179T, 16232A, 16249C, 16304C, 16311C Allentoft 2015; Y-DNA personal communication from author

Mongolia Takhilgat Uzuur-5 [TU17]
Dark brown hair; brown eyes
M 914 BC C M130 A 16223T, 16290T, 16319A, 16362C Hollard 2014

Lithuania Turlojiske [RISE598] M 908-485 BC

H2a2 263G, 15326G Allentoft 2015; Y-DNA personal communication from author
Baltic BA Latvia Kivutkalns [42] F 810-560 cal BCE 585203 H1b1 Mittnik 2017
Baltic BA Latvia Kivutkalns [222] M 805-515 cal BCE 641886 R1a1 U5a1c1 Mittnik 2017
Baltic BA Latvia Kivutkalns [25] M 800-545 cal BCE 682042 R1a1a1b H28a Mittnik 2017
Baltic BA Latvia Kivutkalns [194] M 800-545 cal BCE 130958 R1a1a T1a1b Mittnik 2017
Nordic Late Bronze Age Denmark Trundholm mose II [RISE276] M 794-547 BC 92841 R1b1a2 L265+, PF6500+ T2b 73G, 263G, 709A, 930A, 1438G, 2706G, 3107T, 4917G, 6857T, 8697A, 9861C, 11251G, 11719A, 11812G, 14233G, 14766T1, 4905A, 15326G, 15452A, 15607G, 15928A, 16126C, 16294T, 16296T, 16304C, 16519C Allentoft 2015; Mathieson 2015
Baltic BA Latvia Kivutkalns [215] F 790-535 cal BCE 850417 H1c Mittnik 2017
Baltic BA Latvia Kivutkalns [19] M 730-400 cal BCE 896471 R1a1a1b H10a Mittnik 2017
Baltic BA Latvia Kivutkalns [207] F 730-390 cal BCE 915334 H1b2 Mittnik 2017
Baltic BA Latvia Kivutkalns [209] M 405-230 cal BCE 807138 R1a1a J1b1a1 Mittnik 2017