Iron Age DNA from Europe and West Asia

Mitochondrial and Y-chromosome haplogroups extracted from historic and prehistoric human remains in Europe and related remains in Asia, arranged chronologically. For haplogroup nomenclature and aDNA issues see Introduction to ancient DNA.

Culture

Country

Site and/or Individual

Sex

Date

Genome

Y-DNA

mtDNA

Source



pigmentation from genes
pigmentation surviving

usually approximate number of SNPs Haplogroup Additional information Haplogroup Additional information










Babadag Romania SE near the Black Sea coast [1IA, 2IA]
1000-600 BC


2 samples 16262C incomplete sequences Cardos 2004
Babadag Romania SE near the Black Sea coast [5IA]
1000-600 BC


CRS incomplete Cardoa 2004
Babadag Romania SE near the Black Sea coast [10IA]
1000-600 BC


16192T, 16284C Cardos 2004
Mezocsat Hungary Ludas-Varju-dulo [IR1] Light brown/ dark blond hair, brown eyes M 980-930 BC N 22665262G, 15469724A G2a1 73G, 152C, 263G, 489C, 709A, 750G, 1438G, 2706G, 4769G, 4833G, 5108C, 5601T, 7028T, 7600A, 8547C, 8701G, 8860G, 9377G, 9540C, 9575A, 10398G, 10400T, 10873C, 11719A, 12705T, 13563G, 14200C, 14569A, 14587G, 14766T, 14783C, 15043A, 15301A, 15326G, 16223T, 16278T, 16362C, 16519C Gamba 2014
Iran Tepe Hasanlu, West Azerbaijan (region) [F38] M 971-832 calBCE R1b1a2a2 CTS1078/Z2103 N1a3a Broushaki 2016
Zevakino-Chilikta Kazakhstan Ismailovo [Is1] 900-700 BC D4h1 Unterlander 2017
Zevakino-Chilikta Kazakhstan Ismailovo [Is2] M 900-700 BC 74,469 Q1a F903: 7014317G->C, M1168: 22155597G->A [Additional info Q-L332*] HV-CRS Unterlander 2017; Additional SNPs from Vladimir Tagankin of YFull
Zevakino-Chilikta Kazakhstan Ismailovo [Is3] 900-700 BC H2a1 Unterlander 2017
Zevakino-Chilikta Kazakhstan Zevakino [Ze2] 900-700 BC T2b Unterlander 2017
Zevakino-Chilikta Kazakhstan Zevakino [Ze3] 900-700 BC K1 Unterlander 2017
Zevakino-Chilikta Kazakhstan Zevakino [Ze4, Ze_8] 900-700 BC D4 2 samples Unterlander 2017
Zevakino-Chilikta Kazakhstan Zevakino [Ze5] 900-700 BC I Unterlander 2017
Zevakino-Chilikta Kazakhstan Zevakino [Ze6] F 900-700 BC 163,338 C4 Unterlander 2017
Zevakino-Chilikta Kazakhstan Zevakino [Ze7] 900-700 BC U4a1 Unterlander 2017
Zevakino-Chilikta Kazakhstan Zevakino [Ze9] 900-700 BC D4j3 Unterlander 2017
Thracian Bulgaria Stambolovo [T2G2]
850–700 BC

HV 263G, 750G 1438G 2706G 4769G 7028T 8860G 15326G, 16311C Carpenter 2013

Kazakhstan Zevakinskiy
800-600 BC

H 3 samples CRS Lalueza-Fox 2004

Kazakhstan Zevakinskiy
800-600 BC

U1 16189, 16249 Lalueza-Fox 2004

Kazakhstan Zevakinskiy
800-600 BC

W 16223, 16292 Lalueza-Fox 2004
Thracian Bulgaria Svilengrad [P192-1]
800–500 BC

U3b 750G 1811G 2706G 4640A 4769G 7028T 8860G 9656C 11719A 12308G 12372A 13743C 14139G 15326G 15454C Carpenter 2013

Russia Anashkino hillfort, near Lake Zhizhitsa / Sennitsa, Pskov oblast [A4] M 800-400 BC R1a1
? 16119T, 16137G, 16256T, 16257A 16376C. Reported as H Chekunova 2014
Tagar Russia Novosselovsky region, Anach village, kurgan I, burial 3 [S21]
Blue or green eyes, brown hair, fair or medium skin
? 800 BC–100 AD

T3 16126C, 16189C, 16292T, 16294T Keyser 2009
Tagar Russia Novosselovsky region, Anach village, kurgan I, burial 4 [S22]
Brown eyes
F 800 BC–100 AD

T3 16126C, 16189C, 16292T, 16294T Keyser 2009
Tagar Russia Tchernogorsk, burial 1
Khakassia Republic [S23]
Blue or green eyes
? 800 BC–100 AD

T3 16126C, 16189C, 16292T, 16294T, 16296T Keyser 2009
Tagar Russia Tchernogorsk, burial 6 [S24]
Blue or green eyes
M 800 BC–100 AD R1a1a M17; Riii in table 2 I4 16129A, 16223T, 16304C, 16391A Keyser 2009
Tagar Russia Oust-Abakansty, Khakassia Republic [S25]
Blue or green eyes
M 800 BC–100 AD R1a1a M17; Riv in table 2 G2a 16093C, 16223T, 16234T, 16288C, 16298C, 16327T Keyser 2009
Tagar Russia Beysky region, burial 3
Khakassia Republic [S26] Brown eyes, blond or light brown hair, fair or medium skin
M 800 BC–100 AD R1a1a M17; Rv in table 2 C 16148T, 16223T, 16234T, 16288C, 16298C, 16327T Keyser 2009
Tagar Russia Bogratsky region, kurgan 133, burial 3, Khakassia Republic [S27] F 800 BC–100 AD

H or U CRS Keyser 2009
Tagar Russia Bogratsky region, kurgan 133, burial 3 [S28]
Watson in Table 3
M 800 BC–100 AD R1a1a M17; Rvi in table 2 F1b 16172C,  16179T, 16183C, 16189C,  16232A,  1649C, 604C,  16311C Keyser 2009
Tagar Russia Bogratsky region, kurgan 133, burial 3 [S29] Blue or green eyes M 800 BC–100 AD R1a1a M17; Rvii in table 2 H  or U CRS Keyser 2009
Tagar Russia Bogratsky region, Abakano-Pérévoz II,
burial 1 [S32] Brown eyes
M 800 BC–100 AD R1a1a M17; Rviii in table 2 H5 16304, 16319 Keyser 2009
Aldy Bel Tuva, Russian Federation Arzhan 2 [A2] 700-500 BC A8 Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A3 + A11] 700-500 BC G Two samples Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A4] 700-500 BC D4b1a2a Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A5] 700-500 BC C5b1 Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A6] 700-500 BC U5a1f1 Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A7] 700-500 BC Y1 Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A8] 700-500 BC U4a3 Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A9] 700-500 BC U5a1d2b Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A10 / I0577] M 700-500 BC R1a1a1b S441: 7683058G->A; [Additional info: roughly R-YP1456* (but Y29558-) ] A Unterlander 2017; Additional SNPs from Vladimir Tagankin of YFull
Aldy Bel Tuva, Russian Federation Arzhan 2 [A14] 700-500 BC T1a Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A17] 700-500 BC H Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A19] 700-500 BC C4 Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A20] 700-500 BC G2a Unterlander 2017
Aldy Bel Tuva, Russian Federation Arzhan 2 [A21] 700-500 BC A4 Unterlander 2017
Hallstatt Austria Mitterkirchen [HÜ-I/8] M 700 BC G2a? (Using Whit Athey's haplogroup predictor) STRs only reported

Kiesslich 2005
Nuragic Italy Alghero, Sardinia [AL07]
700 BC

H1? 16189C Caramelli 2007; Ghirotto 2010
Scythian Ukraine Starosillya [SCY006] F 700-600 BCE D4j2 Juras March 2017
Scythian Ukraine Starosillya [SCY009] M 700-600 BCE J2b1a6 Juras March 2017
Scythian Ukraine Starosillya [SCY010] F 700-600 BCE N1b1a Juras March 2017

Kazakhstan Molaly (Kourgan 14,18,20,25)
700-400 BC

M4 16129, 16174, 16223 Lalueza-Fox 2004

Kazakhstan Molaly (Kourgan 14,18,20,25)
700-400 BC

T* 16126, 16294, 16296 Lalueza-Fox 2004
Xiongnu a.k.a. Hsiung-nu? Kazakhstan Birlik (Kourgan 15)
700-200 BC

A* 2 samples 16223, 16319, 16362 Lalueza-Fox 2004
Xiongnu a.k.a. Hsiung-nu? Kazakhstan Birlik (Kourgan 15)
700-200 BC

A* 16223, 16290, 16319, 16362 Lalueza-Fox 2004

Kazakhstan Birlik (Kourgan 15)
700-200 BC

M* 16223, 16286 Lalueza-Fox 2004

Kazakhstan Birlik (Kourgan 15)
700-200 BC

H 16266, 16287 Lalueza-Fox 2004

Kazakhstan Birlik (Kourgan 15)
700-200 BC

U* 16189 Lalueza-Fox 2004
Iberian Spain Puig de la Nau, Benicarlo (Castellon) [PB14, PB20]
700-500 BC

V 2 samples 16293T, 16298C Fernández 2006; Arroyo 2006; Gamba 2008
Iberian Spain Puig de la Nau, Benicarlo (Castellon) [PB18, PB19]
700-500 BC

H 2 samples CRS Gamba 2008
Iberian Spain Castellet de Bernabe, Lıria (Valencia) [CB2]



T1 16163G, 16186T, 16189C, 16250Y, 16294T, 16301Y Gamba 2008
Iberian Spain Los Villares, Caudete de las Fuentes (Castellon) [LV1]



V 16298C Gamba 2008
Iberian Spain Torrelo Bonerot, Castellon [TORR1]



W 16223T, 16292Y, 16295Y, 16304Y, 16311Y Gamba 2008
Iberian Spain Torrelo Bonerot,Castellon [TORR2]



W 16223T, 16292Y, 16295Y, 16304Y Gamba 2008
Iberian Spain Tossal de Sant Miquel, Lıria (Valencia) [TSM1]



H? CRS Gamba 2008
Iberian Spain Tossal de Sant Miquel, Lıria (Valencia) [TSM1]



? 16224Y, 16234Y, 16311Y, reported as K Gamba 2008
Iberian Spain Mas Castellar (Pontos), Girona, Catalonia [1000012-2] ? 700-100 BC

J* −13704 BstNI, 3010G, 16126, 16189 Sampietro 2005

England Heslington, Yorkshire
673-482 BC

J1d
O'Connor 2011
Iberian Spain Illa d’en Reixac (Ullastret), Girona, Catalonia [UE 15112] M 600 BC

U5a 16256, 16270 Sampietro 2005
Iberian Spain Illa d’en Reixac (Ullastret), Girona, Catalonia [UE 1070] M 600 BC

H −7025 AluI, 7028C, 16126, 16311, Sampietro 2005
Iberian Spain Illa d’en Reixac (Ullastret), Girona, Catalonia [15091] ? 600 BC

H1 −7025 AluI, 3010A; CRS in HVI Sampietro 2005
Iberian Spain Illa d’en Reixac (Ullastret), Girona, Catalonia [UE 15224] M 600 BC

H 7028C, 3010G, 13708G, 16126 Sampietro 2005
Iberian Spain Illa d’en Reixac (Ullastret), Girona, Catalonia [UE 15078-1]
Possible decapitation.
M 600 BC

K* 16224, 16311 Sampietro 2005
Scythian Russia Rostov-on-Don [RD7]
600-500 BC

T2 16078G, 16126C, 16294T, 16296T Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD5]
600-400 BC

U5a 16192T, 16256T, 16263C, 16270T, 16399G Der Sarkissian 2011
Iberian Spain Puig de Sant Andreu (Ullastret), Girona, Catalonia [UE 14015] ? 535-200 BC

H 16287 Sampietro 2005
Iberian Spain Puig de Sant Andreu (Ullastret), Girona, Catalonia [UE 14045] ? 535-200 BC

U4 7028T, 12308G, 3010G, 16356 Sampietro 2005
Iberian
Spain
Puig de Sant Andreu (Ullastret), Girona, Catalonia [N.I. 3967 + 1401]
? +M
535-200 BC


H
2 samples 7028C, 3010G, 13708G, 16126 Sampietro 2005
Iberian
Spain
Puig de Sant Andreu (Ullastret), Girona, Catalonia [UE 14020]
?
535-200 BC


T
16126, 16294
Sampietro 2005
Iberian
Spain
Puig de Sant Andreu (Ullastret), Girona, Catalonia [T.C. 4 E I-II]
?
535-200 BC


H1
−7025 AluI, 3010A, 16126 Sampietro 2005
Iberian
Spain
Puig de Sant Andreu (Ullastret), Girona, Catalonia [UE 14015-1]
M
535-200 BC


R0
16362 7028T, 14766C. Recorded as Pre-HV
Sampietro 2005
Iberian
Spain
Puig de Sant Andreu (Ullastret), Girona, Catalonia [UE 14119]
?
535-200 BC



H
16273 −7025 AluI
Sampietro 2005
Iberian
Spain
Puig de Sant Andreu (Ullastret), Girona, Catalonia [2002 Z13]
?
535-200 BC



U5*
16270 16281
Sampietro 2005
Iberian Spain Puig de Sant Andreu (Ullastret), Girona, Catalonia [3613] M 535-200 BC

H 16126, 16311, −7025 AluI, 7028C, 3010G, 13708G Sampietro 2005
Iberian Spain Puig de Sant Andreu (Ullastret), Girona, Catalonia [3615] M 535-200 BC

J* 16126, 13708A, 7028T
Sampietro 2005

Italy Latsch, South Tyrol
500 BC

H90
Brotherton 2013
Early Sarmatian Russia Pokrovka [Pr1] 500-200 BC U3 Unterlander 2017
Early Sarmatian Russia Pokrovka [PR3 /I0575] M 500-200 BC R1b1a2a2 CTS1078:7186135G->C, S20902:18383837C->T [Additional info roughly R-Y21707* (but A12363-)] M Unterlander 2017; Additional SNPs from Vladimir Tagankin of YFull
Early Sarmatian Russia Pokrovka [PR4] 500-200 BC U1a'c Unterlander 2017
Early Sarmatian Russia Pokrovka [PR5] 500-200 BC T Unterlander 2017
Early Sarmatian Russia Pokrovka [PR6] 500-200 BC Fb1 Unterlander 2017
Early Sarmatian Russia Pokrovka [PR7] 500-200 BC N1a1a1a1a Unterlander 2017
Early Sarmatian Russia Pokrovka [PR8] 500-200 BC T2 Unterlander 2017
Early Sarmatian Russia Pokrovka [PR9] 500-200 BC U2e2 Unterlander 2017
Early Sarmatian Russia Pokrovka [PR10] 500-200 BC H2a1f Unterlander 2017
Early Sarmatian Russia Pokrovka [PR11] 500-200 BC T1a Unterlander 2017
Early Sarmatian Russia Pokrovka [PR13] 500-200 BC U5a1d2b Unterlander 2017
Scythian Russia Rostov-on-Don [RD1]
400-300 BC

F1b 16189C, 16232A, 16249, 16304C, 16311C Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD2]
400-300 BC

C 16223T, 16258G, 16298c, 16327T Der Sarkissian 2011
Sarmatian Russia Rostov-on-Don [RD3]



U7 16256T, 16318T Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD6, RD11]



T1a 2 samples 16126C, 16163G, 16186T, 16189C, 16294T Der Sarkissian 2011
Donghu China Jinggouzi, Inner Mongolia
500 BC C* 12 males

M216

D, D4, D5, G2, C and M10 26 samples in all of which 13 were D4, 5 were G2, 5 were C and the rest were single examples. Wang 2012

Kazakhstan Solyanka II (Kourgan 2,3)
500-0 BC

H 16150,  16189 Lalueza-Fox 2004

Kazakhstan Solyanka II (Kourgan 2,3)
500-0 BC

HV 16129 Lalueza-Fox 2004

Kazakhstan Solyanka II (Kourgan 2,3)
500-0 BC

U* 16311 Lalueza-Fox 2004
Scythian Russia Rostov-on-Don [RD8]
500-300 BC

A4 16223T, 16290T, 16319A, 16362C Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD14, RD15]
500-200 BC

I3 2 samples 16086C, 16129A, 16223T, 16391A Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD9]



H rCRS Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD10]



H2a1 16354T Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD12]



U2e 16051G, 16129C, 16189C, 16362C Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD13]
400-300 BC

D 16223T, 16362C Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD16]
400-200 BC

U5a 16192T, 16256T, 16270T, 16311C Der Sarkissian 2011
Scythian Russia Rostov-on-Don [RD17]
400-300 BC

D4b1 16223T, 16239T, 16243C, 16319A, 16362C Der Sarkissian 2011
Sargat Russia Sopinsky, kurgan burial
500 BC-500 AD

T1 16126C, 16163G, 16186T, 16189C, 16243C, 16249T Bennett and Kaestle 2010
Sargat Russia Sopinsky, flat grave burial
500 BC-500 AD

Z 16129A, 16185T, 16215G, 16223T, 16224Y, 16260T, 16298C Bennett and Kaestle 2010
Sargat Russia Kurtuguz I [K2]
500 BC-500 AD

A 16223T, 16290T, 16319A, 16335G Bennett and Kaestle 2010
Sargat Russia Kurtuguz I [K3T]
500 BC-500 AD

C 16223T,16298C, 16327T Bennett and Kaestle 2010
Sargat Russia Kurtuguz I [K3R]
500 BC-500 AD

Z 16093C, 16185T,16223T, 16260T, 16298C Bennett and Kaestle 2010
Scytho-Siberian Altaï Republic Kizil M 500 BC

N1a 16147A, 16172C, 16223T, 16248T,  16355T Ricaut 2004a
Pazyryk Altaï Republic Sebÿstei Valley [SEB 96 K1] M 450 BC

F2a 16093C 16203G 16231C 16291T 16304C  Ricaut 2004c
Pazyryk Altaï Republic
Sebÿstei Valley [SEB 96K2] M 450 BC R1a1a M17
D?
16093C 16129A 16173T 16223T 16319A 16362C Ricaut 2004c; Keyser 2009
Pazyryk Altaï Republic Ak-Alakha-5, mound 5, Ukok Plateau M 400-200 BC

A 16223, 16242, 16290, 16319 Chikisheva 2007
Pazyryk Mongolia Tsengel Khairkhan [TSK07.T1] M


A 16093, 16223, 16242, 16278, 16290, 16311, 16319 Gonzalez-Ruiz 2012
Pazyryk Altaï Republic Ak-Alakha-1, mound 1, burials 1 and 2, Ukok Plateau M 400-200 BC N
Burial 2 (tattooed teenager) was originally thought to be female.
C 16093, 16129, 16223, 16298, 16327 Chikisheva 2007; Pilipenko 2015
Pazyryk Mongolia Baga Turgen Gol [BTG06.T13] M


C 16093, 16129, 16223, 16298, 16327, 16362 Gonzalez-Ruiz 2012
Pazyryk Kazakh Altai Berel' [Be2 ] 400-300 BC D4 Unterlander 2017
Pazyryk Kazakh Altai Berel' [Be3] 400-300 BC H-CRS Unterlander 2017
Pazyryk Kazakh Altai Berel' [Be4] 400-300 BC A Unterlander 2017
Pazyryk Kazakh Altai Berel' [Be6 + Be14] 400-300 BC A6 Unterlander 2017
Pazyryk Kazakh Altai Berel' [Be8] 400-300 BC D4 Unterlander 2017
Pazyryk Kazakh Altai Berel' [Be9] 400-300 BC A4f Unterlander 2017
Pazyryk Kazakh Altai Berel' [Be11 / I0563] M 400-300 BC R1a1a1b2 Z93:7552356G->A [Additional info: R-S23201 +] C4a1+16129 Unterlander 2017; Additional SNPs from Vladimir Tagankin of YFull
Pazyryk Kazakh Altai Berel' [Be12] 400-300 BC HV2 Unterlander 2017
Pazyryk Kazakh Altai Berel' [Be] 400-300 BC A6 Unterlander 2017
Pazyryk Altaï Republic Ak-Alakha-5, mound 3, burial 1, Ukok Plateau M 400-200 BC

D 16223, 16239, 16319, 16362 Chikisheva 2007
Pazyryk Altaï Republic Moinak-2, mound 2, burial 1, Ukok Plateau M 400-200 BC

D 16223, 16311, 16316, 16362 Chikisheva 2007
Pazyryk Mongolia Baga Turgen Gol [BTG05.T1]



D 16223, 16319, 16362 Gonzalez-Ruiz 2012
Pazyryk Mongolia Baga Turgen Gol [BTG05.T8.3, BTG06.T8]



D 2 samples 16223, 16239, 16243, 16319, 16362 Gonzalez-Ruiz 2012
Pazyryk Mongolia Tsengel Khairkhan [TSK07.T2A]



G2 16223, 16227, 16278, 16362 Gonzalez-Ruiz 2012
Pazyryk Altaï Republic Kuturguntas-1, mound 1, Ukok Plateau M 400-200 BC

H? CRS Chikisheva 2007
Pazyryk Altaï Republic Ulandryk-1, mound 14, burial 2, Ulandryk valley M 400-200 BC

H? CRS Chikisheva 2007
Pazyryk Mongolia Olon-Kurin-Gol Valley cemetery 6 [OKG-6-2, 3] F + inf 400-200 BC

HV2 2 samples in same grave of 16217C, 00072C, 00073G, 00152C, +7025 HinfI, -14766 MseI Pilipenko 2010
Pazyryk Mongolia Baga Turgen Gol [BTG06.T10A] M


HV6 16172, 16311 Gonzalez-Ruiz 2012
Pazyryk Mongolia Baga Turgen Gol [BTG06.T3]



J 16069, 16126 Gonzalez-Ruiz 2012
Pazyryk Mongolia Baga Turgen Gol [BTG05.T2, BTG06.T11A, BTG06.T11B]



K 3 samples 16093, 16224, 16311, 16319 Gonzalez-Ruiz 2012
Pazyryk Mongolia Tsengel Khairkhan [TSK07.T2B] M


T1 16126, 16163, 16186, 16189, 16294 Gonzalez-Ruiz 2012
Pazyryk Altaï Republic Ak-Alakha-5, mound 4, burial 1, Ukok Plateau M 400-200 BC

U5 or J 16129, 16182C, 16183C, 16189, 16362 Chikisheva 2007
Pazyryk Mongolia Olon-Kurin-Gol Valley cemetery 10 [OKG-10-1] M 400-200 BC

U5a1 16086C, 16189C, 16192T, 16256T, 16270T, 16304C, 16399G, 00073G, +12308 HinfI. Pilipenko 2010
Pazyryk Altai Republic Verkh-Kaldzhin II M


U5a1 16192T, 16256T, 16270T, 16304C Pilipenko 2010 from Molodin 2003
Pazyryk Mongolia Baga Turgen Gol [BTG05.T8.1, BTG06.T12]



U5a1 2 samples 16192, 16256, 16270, 16304, 16399 Gonzalez-Ruiz 2012
La Tène Germany Glauberg [T1/4 and KL 116-1]
400 BC

K 2 samples 73G 146C 152C 263G 315.1C 16224C 16311C Knipper 2014
La Tène Germany Glauberg [KL 112-1] M 400 BC

I1a 16129A 16172C 16223T 16311C 16391A Knipper 2014
La Tène Germany Glauberg [KL 113-1] F 400 BC

H 263G 315.1C rCRS in HVRI Knipper 2014
La Tène Germany Glauberg [KL 113-2]
400 BC

V 16218T 16298C Knipper 2014
La Tène Germany Glauberg [KL 113-3] F 400 BC

K1a 73G 195C 263G 315.1C 16224C 16311C Knipper 2014
La Tène Germany Glauberg [KL 113-5]
400 BC

H14a 16256T 16352C Knipper 2014
La Tène Germany Glauberg [KL 113-6]
400 BC

H11a2+ 16092C 16293G 16311C Knipper 2014
La Tène Germany Glauberg [KL 116-4]
400 BC

HV + 16311C Knipper 2014
La Tène Germany Glauberg [KL 116-5]
400 BC

U5a1 16192T 16256T 16270T 16399G Knipper 2014

Turkey Ballicaoluk [BOM2]
400-300 BC

H3c C27G, C122G, A123G, C166A, G171A,
A175C, T177d, A181T, T195C, C198T,
T223C, A227C, G229d, G247C, C253T,
A259C, G260C, A263C, A263G, A270C,
A272C, A274C, A286C, A297C, A300C,
C309CCCT, T310C, T455TT, A472C, A479d,
G513C, A515T, A517T, A519C, G564T,
A581T, G611T, A16206G, A16210AG,
C16222CG, C16248CG, T16372TA
Altınışık 2015
Scythian Ukraine Svetlovodsk [SCY001] 400-300 BCE U5a1b Juras March 2017
Scythian Ukraine Svetlovodsk [SCY002] 400-300 BCE J1c2 Juras March 2017
Scythian Ukraine Nesterivka [SCY011] F 400-300 BCE A Juras March 2017
Scythian Ukraine Vapnyarka [SCY012] 400-200 BCE U5b2a1a2 Juras March 2017
Scythian Moldova Glinoe [SCY192] F 400-100 BCE H8c Juras March 2017
Scythian Moldova Glinoe [SCY193] M 400-100 BCE U5a2a1 Juras March 2017
Scythian Moldova Glinoe [SCY196] 400-100 BCE W3a Juras March 2017
Scythian Moldova Glinoe [SCY197] M 400-100 BCE U5a1a1 Juras March 2017
Scythian Moldova Glinoe [SCY300] F 400-100 BCE H5b Juras March 2017
Scythian Moldova Glinoe [SCY303] F 400-100 BCE U5a1a2b Juras March 2017
Scythian Moldova Glinoe [SCY305] 400-100 BCE U5a2b Juras March 2017
Scythian Moldova Glinoe [SCY308] 400-100 BCE F1d Juras March 2017
Scythian Moldova Glinoe [SCY311] F 400-100 BCE T2b Juras March 2017
Scythian Moldova Glinoe [SCY332] F 400-100 BCE M10a1a1a Juras March 2017
Scythian Moldova Glinoe [SCY334] 400-100 BCE H5b Juras March 2017

Russia Sabinka 2 [RISE492] M 396-209 BC R1a
D4b1a2a2 73G, 207A, 263G, 1438G, 3010A, 3107C, 4883T, 8020A, 8414T, 8701G, 9540C, 10181T, 10398G, 10400T, 10873C, 11719A, 12705T, 13720T, 14668T, 14766T, 14783C, 14815T, 14927G, 15043A, 15244G, 15301A. 15326G, 15440C, 15951G, 16223T, 16319A, 16362C Allentoft 2015; Y-DNA personal communication from author
Scythian Russia Nadezhdinka, Volga Steppes, Samara [I0247 / SVP 56] 375-203 calBCE (2220±30 BP) 951,695 R1a1a1b2a2a Z2123 G2a4 Mathieson 2015; Lazaridis 2016

England Linton [L] F 360-50 BC

H1e
Schiffels 2016
Saka Kazakhstan Berel' [BeK11S1] M 350-300 BC


CRS Clisson 2002; Unterlander 2017
Saka Kazakhstan Berel' [BeK11S2] F 350-300 BC

D4g1 16223T, 16362C Clisson 2002; Unterlander 2017

Turkey Karamttepe [KM17]
325-300 BC

H7
C27G, T60TT, C64T, C110G, T115A, C122G, A123G, T125A, C132G, T134G, T142G, T157G, T159G, T161G, C162G, T168G, G171A, T172G, C174G, T177G, G228A, A259G, A263G, T265G, A281G, T282G, A284G, C285G, A291T, A291d, A297C, C299A, A301C, A302C, C315CC, A326T, A328T, C332T, A335G, A336G, A337G, C345T, G347T, A359C, A360C, A385G, A388T, A390G, A396T, G409T, G410T, G412T, G413T, T416d, A446G, T449C,
T450C, A451d, A470G, T471G, T474C,
C483T, A484T, C527T, A565T, A599G,
C628G, A648G, T655A, T656G, G709A,
A750G, G16129A, T16231G, C16292T,
C16294T, C16296T, T16362C, C16408G,
C16549G
Altınışık 2015
Scythian Ukraine Simferopol [SCY005] F 300-100 BCE H Juras March 2017
Przeworsk Poland Gąski [23234]
200 BC-100AD

H5 7028C 16129A, 16304C, reported as H Juras 2012
Przeworsk Poland Gąski [58]
200 BC-100AD

U5a 16256T, 16263C, 16270T Juras 2012
England Melton, East Yorkshire [M1489] F 210 BC-40 AD U2e1e Martiniano 2016

England Hinxton [HI 2] M 170 BC- 80 AD R1b1a2a1a2c1 CTS241/DF13/S521+ H1ag1
Schiffels 2016

England Hinxton [HI 1] M 160 BC-26 AD R1b1a2a1a2c

L21/M529/S145+, S461/Z290+ K1a1b1b Schiffels 2016
Denmark Bøgebjerggård [B4 and B6] M 0 BC/AD H 2 samples 7028C CRS Melchior 2008a

Denmark Bøgebjerggård [B1] F 0 BC/AD

I 7028T, 10034C, 16129A, 16223T, 16391A Melchior 2008a

Denmark Bøgebjerggård [B5] F 0 BC/AD

I 7028T, 10034C, 16129A, 16223T, 16304C, 16391A Melchior 2008a

Denmark Bøgebjerggård [B2] M 0 BC/AD

R0a 7028T, 16126C, 16355T, 16362C Melchior 2008a

Denmark Bøgebjerggård [B3] M 0 BC/AD

U2e 7028T, 12308G, 16129C, 16183C, 16189C, 16362C Melchior 2008a

Denmark Bøgebjerggård [B7] M 0 BC/AD

U5b1 7028T, 12308G, 16074G, 16189C, 16192T, 16249C, 16270T Melchior 2008a
Wielbark Poland Kowalewko [267]
0-300 AD

H5 7028C, 16304C, reported as H Juras 2012
Wielbark Poland Kowalewko [site 12, no. 1]
0-300 AD

H 7028C, CRS Juras 2012
Wielbark Poland Kowalewko [466, 188]
0-300 AD

W 2 samples 8251A, 16223T, 16292T Juras 2012
Wielbark Poland Kowalewko [189]
0-300 AD

W 8251A, 16192T, 16223T, 16292T Juras 2012
Wielbark Poland Kowalewko [314]
0-300 AD

U5b 7768G 16192T, 16270T Juras 2012
Wielbark Poland Kowalewko [502]
0-300 AD

U3 12308G, 14139G, 16343G Juras 2012
Wielbark Poland Rogowo [29 28/99, 680 no. 52/00, 96 13/99, 11/99 object 24, 665 no. 43/00]
100-200 AD

H 5 samples 7028C, CRS Juras 2012
Wielbark Poland Rogowo [718 no. 77/00]
100-200 AD

H 7028C, 16093C, 16129A, 16316G Juras 2012
Wielbark Poland Rogowo [724 no. 80/00]
100-200 AD

H5 7028C, 16153A, 16304C, reported as H Juras 2012
Wielbark Poland Rogowo [438 (grave 68) no. 78/K/99; 45 16/99]
100-200 AD

H 2 samples of 7028C, 16183C, 16189C Juras 2012
Wielbark Poland Rogowo [421 no. 68/K/99]
100-200 AD

H 7028C, 16129A Juras 2012
Wielbark Poland Rogowo [37 no. 32/99]
100-200 AD

J2a 10398G 16069T, 16126C, 16145A, 16231C, 16261T, 16299G Juras 2012
Wielbark Poland Rogowo [705 no. 74/00]
100-200 AD

T2b 16126C, 16294T, 16296T, 16304C Juras 2012
Sarmatian Hungary Csongrád Bokrospuszta 20
0-400 AD

N9a 16223T, 16257A, 16261T Guba 2011; Banffy 2012
Sarmatian Hungary Szakmár 8



N9a 16223T, 16257A, 16261T Guba 2011; Banffy 2012
Sarmatian Hungary Szarvas 23/22



Novel 16235G, 16261T, 16291T, 16293G, 16304C Guba 2011; Banffy 2012
Sarmatian Russia Beslan, Republic of North Ossetia–Alania [A80303] M 100-300 AD J1 M267+ H1c21 Afanasiev, Wen 2015
Sarmatian Russia Beslan, Republic of North Ossetia–Alania [A80304] M 100-300 AD J1 M267+ K1a3 Afanasiev, Wen 2015

Denmark Simonsborg [Si2]
100-200 AD

H 7028C, 16189C Melchior 2010

Denmark Simonsborg [Si4]
100-200 AD

H 7028C, 16172C, 16311C Melchior 2010

Denmark Simonsborg [Si6]
100-200 AD

H 7028C, 16093C, 16221T Melchior 2010

Denmark Simonsborg [Si5]
100-200 AD

I 7028T, 10034C, 16129A, 16223T, 16391A Melchior 2010

Denmark Simonsborg [Si8]
100-200 AD

U5b 7028T, 12308G, 16192T, 16270T, 16304C Melchior 2010

Denmark Simonsborg [Si9]
100-200 AD

T2b 7028T, 15607G, 16126C, 16294T, 16296T, 16304C, 16362C Melchior 2010

Montenegro Velika Gruda [RISE596] F


X1'2'3 153G, 195C, 263G, 338T, 750G, 1438G, 2706G, 3107C, 3531A, 6221C, 6371T, 9250T, 10920T, 11380G, 11719A, 12705T, 13950T, 13966G, 14470C, 14766T, 15326G, 15903G, 15927A, 16223T, 16278T, 16519C Allentoft 2015

Russia Verh-Uimon [RISE600] M
Q1a Q1a1b (M25) K2a5 73G, 146C, 152C, 263G, 280T, 324T, 709A, 750G, 1438G, 1811G, 2706G, 2831A, 3107T, 3480G, 9055A, 9698C, 9716C, 10550G, 11151T, 11299C, 11467G, 11719A, 12308G, 12372A, 14167T, 14766T, 14798C, 15326G, 16224C, 16311C, 16519C Allentoft 2015; Y-DNA personal communication from author, additional info from Felix Emmanuel

Russia Verh-Uimon [RISE601] M
Q1a
M8a1 73G, 263G, 489C, 750G, 1438G, 2706G, 3107T, 4715G, 6179A, 6248C, 7196A, 8453G, 8684T, 9540C, 10398G, 10400T, 10873C, 11719A, 12705T, 14470C, 14766T, 14783C, 15043A, 15047A, 15148A, 15301A, 15326G, 15355A, 15487T, 15697C, 16223T, 16298C, 16319A Allentoft 2015; Y-DNA personal communication from author,

Russia Sary-Bel [RISE602] M
J2a J2a2-PH3085,SK1403 on J2 Research tree: C4+152 73G, 152C, 263G, 489C, 750G, 1438G, 2706G, 3107T, 3552A, 4065G, 4742C, 7196A, 8701G, 9540C, 9545G, 10398G, 10400T, 10873C, 11719A, 11914A, 11969A, 12705T, 13263G, 13641C, 14318C, 14766T, 14783C, 15043A, 15204C, 15301A, 15326G, 15487T, 16223T, 16297C, 16298C, 16327T, 16519C Allentoft 2015; Y-DNA pers. comm. from author + additional info from Chris Rоttensteiner
Tachtyk Russia Bogratsky region, Abakano-Pérévoz I,
burial 5, Khakassia Republic [S33] Blue or green eyes, blond or light brown hair, fair or medium skin
F 100–400 AD

C 16093C, 16129A, 16223T, 16298C, 16327T Keyser 2009
Tachtyk Russia Bogratsky region, Abakano-Pérévoz I,
burial 4 [S34] Blue or green eyes
M 100–400 AD R1a1a M17; Riii in table 2 HV 16172C, 16311C Keyser 2009
Tachtyk Russia Bogratsky region, Abakano-Pérévoz I,
burial 2 [S35] Brown eyes
F 100–400 AD

H or U 16093C, 16209C Keyser 2009
Tachtyk Russia Bogratsky region, Abakano-Pérévoz I, burial [S36]
Blue or green eyes, blond or light brown hair, fair or medium skin
F 100–400 AD

N9a 16223T, 16257A, 16261T Keyser 2009
Tachtyk Russia Bogratsky region, Abakano-Pérévoz I, burial 4 [S37]
Blue or green eyes, blond or light brown hair, fair or medium skin
F 100–400 AD

T1 16126C, 16163G, 16186T, 16189C, 16269G, 16294T, 16362C Keyser 2009
Przeworsk Poland Karczyn [353 PET 30/580]
100-500 AD

H 7028C, 16222T Juras 2012
Przeworsk Poland Karczyn [PET 30/148 ob. 157]
100-500 AD

N1a1 10238C, 16147A, 16223T, 16248T, 16320T, 16355T Juras 2012

Denmark Skovgaarde [S4] ? 200-270 AD

H 7028C, 16311C Melchior 2008a

Denmark Skovgaarde [S5] F 200-270 AD

H 7028C, 16162G, 16266T, 16319A Melchior 2008a

Denmark Skovgaarde [S6] F 200-270 AD

H 7028C, 16299G Melchior 2008a

Denmark Skovgaarde [S14] F 200-270 AD

H 7028C, 16263C, 16319A Melchior 2008a

Denmark Skovgaarde [S3] F 200-270 AD

H5 7028C, 16304C, reported as H1 Melchior 2008a

Denmark Skovgaarde [S1] ? 200-270 AD

J 7028T, 13708A, 16069T, 16126C Melchior 2008a

Denmark Skovgaarde [S9]
200-270 AD

J 7028T, 13708A, 16069T, 16093C, 16126C Melchior 2008a

Denmark Skovgaarde [S2] M 200-270 AD

K 7028T, 12308G 16224C, 16311C Melchior 2008a

Denmark Skovgaarde [S11] F? 200-270 AD

K 7028T, 12308G 16093C, 16224C, 16311C Melchior 2008a

Denmark Skovgaarde [S13] F 200-270 AD

U3a 7028T, 12308G 16343G, 16390A Melchior 2008a

Denmark Skovgaarde [S7] F 200-270 AD

V 7028T, 4580A, 16298C Melchior 2008a

Kazakhstan Kok-Mardan (Kourgan 2,3,6,16)
200-500 AD

U1 16183, 16189, 16249 Lalueza-Fox 2004
Alan Russia Republic of North Ossetia–Alania [A80305] M 300-400 AD R1a1a1b2a2 Z94+, Z95+ W1c Afanasiev, Wen 2015
Alan Russia Republic of North Ossetia–Alania [A80307] M 400-500 AD G2a P15+ X2i Afanasiev, Wen 2015

Sweden Oxe 7 [RISE174] M 427-611 AD

W1 73G, 263G, 709A, 750G, 1243C, 1438G, 2706G, 3107T, 3505G, 5460A, 7864T, 9020T, 11053G, 11347G, 11674T, 11719A, 11947G, 12414C, 12705T, 15326G, 16223T, 16292T, 16519C Allentoft 2015

Russia Kytmanovo [RISE504] M 721-889 AD J2 J2a2-PH3085,SK1403

on J2 Research tree:

C4a1d 73G, 195C, 263G, 489C, 1438G, 1715T, 2706G, 3107T, 3552A, 4353C, 7196A, 9545G, 10398G, 10400T, 11719A, 11969A, 12672G, 12705T, 12940A, 13263G, 14318C, 14766T, 14783C, 15043A, 15204C, 15301A, 15326G, 15487T, 15968C, 16129A, 16150T, 16223T, 16298C, 16327T, 16519C Allentoft 2015; Y-DNA personal communication from author + additional info from Chris Rоttensteiner