Mesolithic Western Eurasian DNA

Mitochondrial and Y-chromosome haplogroups extracted from prehistoric hunter-gathers after the Last Glacial Maximum in Europe and related remains in Asia, arranged chronologically. Pitted Ware sites are classed as Mesolithic here, because of their hunter-gatherer economy, though they fall within the Neolithic period. For haplogroup nomenclature and aDNA issues see Introduction to ancient DNA

Culture

Country

Site and/or Individual

Sex

Date

Genome

Y-DNA

mtDNA

Source

Pigmentation from genes usually approximate number of SNPs Haplogroup Additional information Haplogroup Additional information
England, UK Gough's Cave, Cheddar [Not Cheddar Man] 12,700 BC Unreliable CRS. Contamination likely. Sykes 2000; Jacobi 2009 for dating
Natufian

Israel Raqefet Cave, Mount Carmel [I0685 / Nat 4] M 11840-9760 BCE CT CTS9555, Y1580 Lazaridis 2016
Natufian

Israel Raqefet Cave, Mount Carmel [I1069 / Nat 5] M 11840-9760 BCE E1b1 (xE1b1a1, E1b1b1b1) M5403+ (E), P179+ (E1b1), (Z1116-, CTS8649-) Lazaridis 2016
Natufian

Israel Raqefet Cave, Mount Carmel [I1690 / Nat 6] M 11840-9760 BCE CT Y1462, M5723, L977 Lazaridis 2016
Natufian

Israel Raqefet Cave, Mount Carmel [I1072 / Nat 9] M 11840-9760 BCE E1b1b1b2 (xE1b1b1b2a, E1b1b1b2b) CTS8182+, CTS11781+ (E1b1b1b2), (CTS1652-, CTS11051-, CTS11574-) N1b Lazaridis 2016
Natufian

Israel Raqefet Cave, Mount Carmel [I0861 / Nat 10] M 11840-9760 BCE E1b1b1b2 (xE1b1b1b2a, E1b1b1b2b) L336+ (E1b1b), M5108+, CTS3637+, CTS7154+, PF1755+, L796+ (E1b1b1), CTS11781+ (E1b1b1b2), (L857-, Z865-) Lazaridis 2016
Natufian

Israel Raqefet Cave, Mount Carmel [I0687 / Nat 13] F 11520-11110 calBCE (11405±120 BP) Lazaridis 2016
Italy Grotta Continenza, Abruzzo [Continenza] F 11,200-10,510 cal BP U5b1 Fu 2013; Fu 2016
Portugal Unreliable Various claimed haplogroups. Museum bones, contamination likely. Chandler 2005
Germany Blätterhöhle, Hagen [BLA 3] 9210 ± 29 BC U2e 16129C, 16189C, 16362C, 16391A Bollongino 2013
Iran Hotu Cave [I1293 / Hotu IIIb] M 9100-8600 BCE

J (xJ2a1b3, J2b2a1a1)

[J2a-CTS1085]

PF4519+, FGC3271+, PF4530+, CTS5934+, F2839+, PF4619+ (L227-, Z639-) Lazaridis 2016; [additional analysis of haplogroup by Chris Rottensteiner ]
Germany Blätterhöhle [BLA 6] 8796 ± 90 BC U5b2a2 16172C, 16189C, 16270T, 16398A Bollongino 2013
Germany Blätterhöhle [BLA 2] 8748 ± 67 BC U/K 12308G Bollongino 2013
Germany Blätterhöhle [BLA 20] 8652 ± 58 BC U5a2c3* 73G, 263G, 750G, 1438G, 2706G, 3197C, 4769G, 7028T, 8860G, 9477A, 10619T, 11465C, 11467G, 11719A, 12308G, 12372A, 13617C, 14766T, 14793G, 15326G, 16192T, 16256T, 16270T, 16526A, 16192T, 16256T, 16270T Bollongino 2013
Germany Blätterhöhle [BLA 19] 8638 ± 56 BC U5a 16192T, 16256T, 16270T, 16294T 12308G 14798T Bollongino 2013
England, UK Gough's Cave, Cheddar [Cheddar Man] M 8000 BC Unreliable; U5 16192T, 16270T, reported as U5a. Never properly published. Contamination likely. Due to be re-tested. ISOGG; personal communication Chris Stringer re contamination.
France L'abri des Cabônes [Ranchot88] F 10,240-9,930 cal BP U5b1 G3531A,8276.1C, T16189C!, T16192C!, C16519T, (8276.1C) Posth 2016; Fu 2016
France Les Closeaux (Rueil-Malmaison) [LesCloseaux 13] F 10,240-9,560 cal BP U5a2 G54A, G7805A, C16519T Posth 2016; Fu 2016
Georgia Kotias Klde [KK1] M 7940-7600 calBCE [7938-7580 calBCE (8665±65 BP, RTT-5246); 7946-7612 calBCE (8745±40, OxA-28256)]

1,230,084

J2a M410> PF4610> Z6049> SK1321(xSK1314), Y12630+, Y12628+, SK1314/Y12595-, Y12624-, CTS3089-

H13c Mutations with respect to the Reconstructed Sapiens Reference Sequence: 73A, 146T, 152T, 195T, 247G, 769G, 825T, 1018G, 2706A, 2758G, 2885T, 3594C, 4104A, 4312C, 4769A, 7028C, 7146A, 7256C, 7521G, 8206A, 8468C, 8655C, 8701A, 9540T, 10289G, 10398A, 10664C, 10688G, 10810T, 10873T, 10915T, 11719G, 11914G, 12705C, 13105A, 13276A, 13506C, 13650C, 14766C, 14872T, [16129G not found], 16187C, 16189T, 16223C, 16230A, 16278C, 16311T, 16519T Jones 2015; Fu 2016; Lazaridis 2016; additional information on SNPs from Chris Rоttеnѕtеіnеr
Russia Chekalino ? 7800 BC U5a 16192T, 16256T, 16270T, 16294T, +12308 HinfI Bramanti 2009
Russia Lebyazhinka [IV a] M 8000-7000 BC U5a1 16192T, 16241C, 16256T, 16270T, 16399G; +12308 HinfI Bramanti 2009
Greece Theopetra, Thessaly [Theo5] 7605–7529 BC K1c Hofmanová 2015
Greece Theopetra, Thessaly [Theo1] 7288–6771 BC K1c Hofmanová 2015
Germany Falkensteiner Höhle M 9,410-8,990 cal BP F P143, P187 U5a2c A4732G, G8572A, A16171G, C16519T, T16189C! missing Bramanti 2009; Posth 2016; Fu 2016
Portugal Toledo, Lourinha [K-13]
7200-6800 BC R? 16223T Fernández 2005
Sweden Stora Förvar cave, Stora Karlsö Island [SfF11] M 7500-7250 cal. BP U5a1 Skoglund 2014
Sweden Stora Förvar cave, Stora Karlsö Island 6,873 ± 119 cal BC U4b1 Skoglund 2013
Germany Bad Dürrenberg [2] F 6850 BC U4 16356C, +12308 HinfI, +7028 AluI Bramanti 2009
Russia Yuzhnyy Oleni Ostrov [I0061 / UzOO 74; Karelia in Fu 2016] M 6850-6000 BC

1,057,094 R1a1*

M459+, Page65.2+, M515-, M198-, M512-, M514-, L449-

C1g Reported as C1f in Der Sarkissian 2014. C146T, C152T, T182C!, C195T, G247A!, A249d, 290–291d, T489C, 522.AC, A769G, A825t, A1018G, A2758G, C2885T, T3552a, T3594C, G4104A, T4312C, A4715G, G7146A, C7196a, T7256C, A7521G, T8468C, A8577G, G8584A, T8655C, A9545G, C10400T, T10664C, A10688G, C10810T, C10915T, A11605t, G11914A!, A12217G, G13105A, A13263G, G13276A, T13506C, T13650C, T14318C, T14783C, G15043A, G15301A, A15487t, A16129G, A16183c, T16187C, T16189C!, G16230A, T16278C, T16298C, C16311T, T16325C, C16327T, C16519T Der Sarkissian 2011; Der Sarkissian 2013; Der Sarkissian 2014; Haak 2015; Mathieson 2015; Fu 2016; Lazaridis 2016
Germany Hohlenstein-Stadel [5830a-c] M 8,980-8,440 cal BP U5b2c1 C16519T, (960.XC, T14182C) Bramanti 2009; Brandt 2013; Posth 2016; Fu 2016
Italy Grotta d’Oriente, Favignana, Ègadi Islands, Sicily [Oriente B] 8,740-8,390 cal BP HV1? or U5b3d? 16067T. Reported as HV1, but HVR-1 only tested. 16067T also appears in U5b3d. Mannino 2012
Kunda Culture Lithuania Spiginas [4] F 6350 BC U4 16356C, +12308 HinfI, +7028 AluI Bramanti 2009
Kunda Culture? Lithuania Donkalnis [1] F U5b 16192T, 16270T, +12308 HinfI, 14793A Bramanti 2009; Brandt 2013 (updated haplogroup)
Luxembourg Loschbour, Heffingen [LSB 1] dark hair, 50% probability of blue eyes, darker skin than Neolithic sample M 6210-5990 calBCE (7205±50 BP, OxA-7738) 1,240,000 I2a1b L178+, M423+, P37.2+, L460+, M438+, L68+, P38+, M170+, M359.2-, L161.1-, L621- reported as I2a1b* U5b1a T16189C!, A6701G Fu 2013; Delsate 2009; Lazaridis 2013; Haak 2015; Mathieson 2015; Fu 2016; Lazaridis 2016
Kitoi Russia Lokomotiv cemetery, Irkutsk [LOK_1980.012 and LOK_1980.017] M 6125–4885 BC missing data A 2 samples 16223T, 16290T, 16319A Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv cemetery, Irkutsk [OK_1985.031.02] M 6125–4885 BC C3-M217 A 16223T, 16290T, 16319A Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv, Irkutsk [LOK_1980.022.02] M 6125–4885 BC K-M9 C 16223T, 16298C, 16327T Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv, Irkutsk [LOK_1980.004] M 6125–4885 BC missing data D 16223T Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv, Irkutsk [LOK_1980.016] M 6125–4885 BC K-M9 D 16223T Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv, Irkutsk [LOK_1980.006 and OK_1981.024.01] M 6125–4885 BC R1a1-M17 2 samples F 2 samples 16232A, 16249C, 16304C, 16311C Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv, Irkutsk

[LOK_1980.010.03]

M 6125–4885 BC not assigned F 16232A, 16249C, 16304C, 16311C Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv, Irkutsk, [LOK_1988.038.01, LOK_1990.044.01 and LOK_1990.044.02] M 6125–4885 BC missing data 3 samples F 3 samples 16232A, 16249C, 16304C, 16311C Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv, Irkutsk [LOK_1990.042] M 6125–4885 BC K-M9 G2a 16223T, 16227G, 16278T Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv, Irkutsk [LOK_1980.014.03] M 6125–4885 BC missing data U5a 2 samples 16256T, 16270T Mooder 2006; Moussa 2016
Kitoi Russia Lokomotiv, Irkutsk [LOK_1980.010.02] M 6125–4885 BC K-M9 not known Mooder 2006; Moussa 2016
Croatia Vela Spila/Island Korčula M 6080-6020 BC U5b2a5 16270C 16311C A16129G T16187C C16189T T16223C G16230A C16270T T16278C Szécsényi-Nagy 2014
Russia Zhokhov, New Siberian Islands [C130L] 6000 BC UK/V? –9052 HaeII , +12308 HinfI, +10394 DdeI, –9053 HhaI, +4643 RsaI, 16298C, 16319A Pitulko 2015
Russia Zhokhov, New Siberian Islands [C130R2a ] 6000 BC U-K +8249/ AvaII, –9052 HaeII/ +12308 HinfI, +10394 DdeI, –9053 HhaI/ +4643 RsaI, 16093C/t, 16223T/c, 16224T/c, 16293A/g, 16311C, 16319A/g, 16355T/c Pitulko 2015
Russia Zhokhov, New Siberian Islands [ED140, Oo23, Pp240] 6000 BC U-K 3 samples +8249/ AvaII, –9052 HaeII/ +12308 HinfI, +10394 DdeI, –9053 HhaI/ +4643 RsaI, 16093C/t, 16223T/c, 16224T/c, 16293A/g, 16311C, 16319A/g, 16355C/t Pitulko 2015
Russia Zhokhov, New Siberian Islands [Nn200] 6000 BC W? +8249/ AvaII –9052 HaeII/ +12308 HinfI +10394 DdeI –9053 HhaI/ +4643 RsaI, 16016C, 16017C, 16223T, 16292T, 16311C, 16319A Pitulko 2015
Russia Zhokhov, New Siberian Islands [Nn220-1 and Nn220-2] 6000 BC U-K 2 samples –9052 HaeII +12308 HinfI +10394 DdeI –9053 HhaI +4643 RsaI, 16093C, 16224C, 16293G, 16311C, 16355T Pitulko 2015
Poland Janisławice 6000 BC U5b1b 16189C 16270T Stanaszek 2013
Germany Felsdach Inzigofen 8,980-8,380 cal BP U5a2c T152C!,735.1G, 11728T, C16192T, C16519T Posth 2016; Fu 2016
Germany Große Ofnet Höhle F 8,430-8,060 cal BP U5b1d1 C16176T, C16519T, (525insAC) Posth 2016; Fu 2016
France Les Vignolles (Mareuil-lès-Meaux) [MareuilLesMeaux1] 8320±90 BP U5a2 T16362C, C16519T, (6008N) Posth 2016
Germany Groß Fredenwalde [1962 excavation 3] F 6048 ± 22 BC U5b 12308G, 16270T Terberger 2015
Spain La Braña-Arintero, Leon [LB 2] M 7960-7750 cal BP = 5960-5750 BC U5b2c1 As La Braña 1 Sanchez-Quinto 2012
Spain La Braña-Arintero, Leon [LB 1] dark hair, probability of blue eyes, darker skin than Neolithic sample M 5990-5740 calBCE 1,057,168 C1a2 M130, M216, P255, P260, V183, V199, V232, V20

Reported as C6 (nomenclature up to 24 Jan 2014)

U5b2c1 150T, 723G, 960.1C. 1721T, 3197C, 6920A, 7768G, 9477A, 12372A, 13017G, 13434G, 13617C, 13637G, 14182C, 16192T, 16270T Sanchez-Quinto 2012; Olalde 2014; Mathieson 2015; Fu 2016
Sweden Motala [1] F 5898-5531 BC 643,050 U5a1 G5460A Lazaridis 2013; Mathieson 2015
Sweden Motala [2] M 5898-5531 BC 805,855 I2c2 PF3827+, L597+, P38+, U179+ , L41+, M253-, L37- U2e1 C16527T Lazaridis 2013; Haak 2015; Mathieson 2015
Sweden Motola [3] M 5898-5531 BC 320,332 I2a1b* M423+, L68+, M258+, U179+, L181-, L417- U5a1 G5460A, A9389G Lazaridis 2013; Haak 2015; Mathieson 2015
Sweden Motala [4] F 5898-5531 BC 763,001 U5a2d A13158G Lazaridis 2013; Haak 2015; Mathieson 2015
Sweden Motala [6] M 5898-5531 BC 611,053 I2a1* P37.2+, L55+, L232-, L159.1- U5a2d C152T!, G6480A Lazaridis 2013; Haak 2015; Mathieson 2015
Sweden Motala [9] M 5898-5531 BC I2a1a1a* L672+, P38+, P40-, Z118- U5a2 G228A, G1888A, A2246G, C3756T, G6917A, A9531G Lazaridis 2013; Haak 2015
Sweden Motala [12] M 5898-5531 calBC 944,923 I2a1b L147.2+, L178+, M423+, L178+, M423+, P37.2+, L460+, L68+, M170+, M258+, U179+, M359.2-, L621- U2e1 C16527T Lazaridis 2013; Haak 2015; Mathieson 2015; Fu 2016
Germany Groß Fredenwalde [1962 excavation 1] M 5825 ± 48 BC U5b 12308G, 16270T Terberger 2015
Russia Sok River, Samara [I0124/SVP 44] M 5650-5555 BC R1b1a M343+, L278+, [P297 equivalent PF6513+], M478-, [M478 equivalent Y13872+, Y13866- (The presence of positive and negative markers in the M478 node can reflect an intermediate stage of its formation.)], M478-, M269- U5a1d C146T, C152T, C195T, A247G, A249d, 290-291d, T489C, A769G, A825t, A1018G, A2758G, C2885T, T3552a, T3594C, G4104A, T4312C, A4715G, G7146A, C7196a, T7256C, A7521G, T8468C, A8577G, G8584A, T8655C, A9545G, C10400T, T10664C, A10688G, C10810T, C10915T, A11605t, A12217G, G13105A, A13263G, G13276A, T13506C, T13650C, T14318C, T14783C, G15043A, G15301A, A15487t, A16129G, T16187C, G16230A, T16278C, T16298C, C16311T, T16325C, C16327T, C16519T, A8577G, A11605t, A12217G, T16189C! Haak 2015; Sergey Malyshev; Mathieson 2015
Germany Bockstein Höhle F 8,370-8,160 cal BP U5b1d1 C16176T, C16519T, (525insAC) Posth 2016; Fu 2016
France Les Fontinettes, Cuiry-lès-Chaudardes [Chaudardes 1] M 8,360-8,050 cal BP I PF3640, CTS5764, Z16987 U5b1b 6056T, 6057T, T16092C, C16327T, C16519T, (7362N) Posth 2016; Fu 2016
Russia Yuzhnyy Oleni Ostrov [UzOO 43 and 45] 7500 BP U2e 2 samples 16129C, 16189C, 16362C Der Sarkissian 2011; Der Sarkissian 2013
Russia Yuzhnyy Oleni Ostrov [UzOO 16 and 40] 7500 BP U4 2 samples 16093C, 16356C Der Sarkissian 2011; Der Sarkissian 2013
Russia Yuzhnyy Oleni Ostrov [UzOO 70] 7500 BP U5a 19192T, 16256T, 16270T, 16318G Der Sarkissian 2011; Der Sarkissian 2013
Russia Yuzhnyy Oleni Ostrov [UzOO 77] 7500 BP H? 16235G, 16311C, 16362C Der Sarkissian 2011; Der Sarkissian 2013
Russia Yuzhnyy Oleni Ostrov [UzOO 7 and 8] 7500 BP C1g Reported as C1f in Der Sarkissian 2014. 2 samples C146T, C152T, T182C!, C195T, G247A!, A249d, 290–291d, T489C, 522.AC, A769G, A825t, A1018G, A2758G, C2885T, T3552a, T3594C, G4104A, T4312C, A4715G, G7146A, C7196a, T7256C, A7521G, T8468C, A8577G, G8584A, T8655C, A9545G, C10400T, T10664C, A10688G, C10810T, C10915T, A11605t, G11914A!, A12217G, G13105A, A13263G, G13276A, T13506C, T13650C, T14318C, T14783C, G15043A, G15301A, A15487t, A16129G, A16183c, T16187C, T16189C!, G16230A, T16278C, T16298C, C16311T, T16325C, C16327T, C16519T Der Sarkissian 2011; Der Sarkissian 2013; Der Sarkissian 2014; Haak 2015
Russia Yuzhnyy Oleni Ostrov [I0211/UzOO 40] M 5500-5000 BC J PF4521, F2114, CTS5934, CTS7028, CTS7229, FGC1599, YSC0000228, CTS11291 U4a Mathieson 2015
Russia Popova [Po 4] 7000 BP U4 16356C Der Sarkissian 2011; Der Sarkissian 2013
Russia Popova [Po 2] 7000 BP U4 16093C, 16356C Der Sarkissian 2011; Der Sarkissian 2013
Spain La Chora (Cantabria) [CH-1]
5346 ± 90 BC

Date from Strauss 2002



? 73G, 16093, 16362, reported as H6 Hervella 2012
Spain Cingle del Mas Nou, Castellon 5000 BC U5 2 samples 16270T Olaria 2011
Spain Aizpea, Navarre [AIZ-1] 4600 +/- 65 BC U5b1 16051, 16093, 16189, 16192, 1270 Hervella 2012
France Le Vieux Tordoir, Berry-au-Bac [BerryAuBac1] M 7,320-7,170 cal BP I F929, CTS2514, PF3836 U5b1a T16192C!, C16519T Posth 2016; Fu 2016
Narva Culture Lithuania Kretuonas [3] M 4450 BC U5b 16192T, 16270T, +12308 HinfI, 14793A Bramanti 2009; Brandt 2013
Narva Culture Lithuania Kretuonas [1] F 4200 BC U5b 16192T, 16270T, +12308 HinfI, 14793A
Bramanti 2009; Brandt 2013
Zedmar culture? Poland Dudka [2] ? 3650 BC U5b 16189C, 16270T, +12308 HinfI
Bramanti 2009; Brandt 2013
Zedmar Culture Poland Dudka [3] ? 4000-3000 BC U5b 16189C, 16265G, 16270T, +12308 HinfI, +7028 AluI
Bramanti 2009
Russia Serteya (Smolenskaya oblast) VIII [А3] M 4000 BC R1a1 ? 16132T, 16163T, 16166G, 16256T. Reported as H Chekunova 2014
Pitted Ware Sweden Kopingsvik, Oland [GE 33 (KOP 29)] 3200–2850 BC T2b 16126C 16294 T 16296T 16304C Malmstrom 2015
Pitted Ware Sweden Kopingsvik [GE 44 (KOP 32)] 3200-2850 BC H1f 16093C Malmstrom 2015
Pitted Ware Sweden Kopingsvik [GE 44 (KOP 32)] 3200-2850 BC K1a1 16093C 16224C 16311C Malmstrom 2015
Pitted Ware Sweden Kopingsvik [GE 44 (KOP 32)]
3200-2850 BC U4a1 16134T Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 4] 3000-2400 BC U4 16093C 16356C Malmstrom 2015
Pitted Ware Sweden Fridtorp, Västerhejde, Gotland [Fri 24] 3000-2400 BC U4 16093C 16356C Malmstrom 2015
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 4, 5] 3000-2400BC U4 2 samples 16356C Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 29B] 3000-2400 BC HV0 16298C Malmstrom 2015
Pitted Ware Sweden Visby, Gotland [Vis 30B] 3000-2400 BC U5b1 16189C 16192 T 16270T 16362C Malmstrom 2015
Pitted Ware Sweden Fridtorp [Fri 28] 3000-2400 BC K1a1 16093C 16224C 16311C Malmstrom 2015
Pitted Ware Sweden Visby [Vis 32] 3000-2400 BC U5a 16192T 16256 T 16270T Malmstrom 2015
Pitted Ware Sweden Visby [Vis 7B] 3000-2400 BC K1a1 16093C 16224C 16311C Malmstrom 2015
Pitted Ware Sweden 3000-2400 BC Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 14, 19]
2800-2000 BC H/R 2 samples A16129G T16187C C16189T T16223C G16230A T16278C Malmstrom 2009; Brandt 2013
Pitted Ware Sweden Ajvide [Ajv 13] 2800-2000 BC U4 A16129G T16187C C16189T T16223C G16230A C16260T T16278C C16311T T16356C Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 52b, 66] 2800-2000 BC U4 2 samples T16093C A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Brandt 2013
Pitted Ware Sweden Ajvide [Ajv 70] M 2800-2000 BC U4 T16093C A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Skoglund 2012; Brandt 2013; Skoglund 2014
Pitted Ware Sweden Ajvide [Ajv 54 ] 2800-2000 BC U5b A16129G T16187C C16189T T16223C G16230A C16270T T16278C C16311T Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 36] 2800-2000 BC U5b2b1a A16129G T16187C C16189T T16223C G16230A C16270T T16278C C16311T T16362C Malmstrom 2009; Brandt 2013; ; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 5, 29] 2800-2000 BC U5a1a'g 2 samples A16129G T16187C C16189T T16223C G16230A C16256T C16270T T16278C C16311T Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 52A] M 2800-2000 BC HV0 A16129G T16187C C16189T T16223C G16230A T16278C T16298C C16311T Malmstrom 2009; Skoglund 2012; Brandt 2013; Skoglund 2014; Malmstrom 2015
Pitted Ware Sweden Ajvide [Ajv 53] F 2800-2000 BC U4d Skoglund 2014
Pitted Ware Sweden Ajvide [Ajv 58] M 2800-2000 BC I2a1 M429+, P126+, P129+, L460+, P37.2+ U4d Skoglund 2014
Pitted Ware Sweden Ajvide [Ajv 59] M 2800-2000 BC U Skoglund 2014
Pitted Ware Sweden Fridtorp, Västerhejde, Gotland [Fri 15, 22] 2800-2000 BC U4 2 samples A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Fridtorp [Fri 4] 2800-2000 BC U5b A16129G, T16187C, C16189T, T16223C, G16230A, C16270T, T16278C, C16311T, Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Fridtorp [Fri 27] 2800-2000 BC U5a1a'g A16129G, T16187C, C16189T, T16223C, G16230A, C16256T, C16270T, T16278C, C16311T Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 8] M 2800-2000 BC U4d T629C A769G A825t A1018G C1224T C1226T G1315A A1811G G2174A G2182A G2185A A2758G C2885T T3594C G4104A G4153A C4189T T4312C C5211T C5974T T5999C A6047G G6075A C6181T G6755A G6993A G7146A T7256C C7412T C7427T A7521G C7974T C7978T C8351T C8375T T8468C T8655C G8701A G8860A G8998A G9192A C9540T C10088T C10213T G10398A T10664C A10688G C10810T C10873T C10915T C11332T A11337G A11467G A11914G A12308G G12372A T12705C G13105A G13276A T13506C T13650C C13860T C14620T C15049T T15693C T16093C A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Skoglund 2012; Brandt 2013; Skoglund 2014; Malmstrom 2015
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 6B] 2800-2000 BC T2b T16126C A16129G T16187C C16189T T16223C G16230A T16278C C16294T C16296T T16304C C16311T Malmstrom 2009; Brandt 2013
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 9] 2800-2000 BC K1a T16093C A16129G T16187C C16189T T16223C T16224C G16230A C16242T T16278C Malmstrom 2009; Brandt 2013; Malmstrom 2015
Pitted Ware Sweden Ire, Hangvar, Gotland [Ire 3] 2800-2000 BC U4 A16129G T16187C C16189T T16223C G16230A T16278C C16311T T16356C Malmstrom 2009; Brandt 2013; ; Malmstrom 2015
Poland Drestwo [2] M 2250 BC U5a 16192T, 16256T, 16270T, +12308 HinfI Bramanti 2009
Russia Bolshoy Oleni Ostrov [BOO 49-3, 57-1] 3500 BP U4a1 2 samples 16093C, 16129A, 16311C, 16356C Der Sarkissian 2011; Der Sarkissian 2013
Russia Bolshoy Oleni Ostrov [BOO 49-1, 72-11] 3500 BP U5a 2 samples 16192T, 16256T, 16270T Der Sarkissian 2011; Der Sarkissian 2013
Russia Bolshoy Oleni Ostrov [BOO 72-9, 72-10, 72-14, 78-8] 3500 BP U5a1 4 samples 16192T, 16256T, 16270T, 16399G Der Sarkissian 2011; Der Sarkissian 2013
Russia Bolshoy Oleni Ostrov [BOO 72-4] 3500 BP T* 16093C, 16126C, 16294T Der Sarkissian 2011; Der Sarkissian 2013
Russia Bolshoy Oleni Ostrov [BOO 49-2, 49-4, 57-3, 72-2, 72-7, 72-12] 3500 BP C* 6 samples 16223T, 16298C, 16327C Der Sarkissian 2011; Der Sarkissian 2013
Russia Bolshoy Oleni Ostrov [BOO 72-5, 72-6] 3500 BP C5 2 samples 16148T, 16223T, 16288C, 16298C, 16311C, 16327C Der Sarkissian 2011; Der Sarkissian 2013
Russia Bolshoy Oleni Ostrov [BOO 49-6, 72-13, 72-15] 3500 BP D* 3 samples 16223T, 16362C Der Sarkissian 2011; Der Sarkissian 2013
Russia Bolshoy Oleni Ostrov [BOO 49-5, 72-3] 3500 BP Z1a 2 samples 16129A, 16185T, 16223T, 16224C, 16260T, 16298C Der Sarkissian 2011; Der Sarkissian 2013
Russia Bolshoy Oleni Ostrov [BOO 72-1] 3500 BP Z1a 16129A, 16155G, 16185T, 16223T, 16224C, 16260T, 16298C Der Sarkissian 2011; Der Sarkissian 2013